Part 43

We have now to note several species of umbelliferous plants that grow in bogs and other wet places. The general features of the order (Umbelliferæ) will be found on p. 167, and the reader should refer to these, if necessary, before attempting to identify the following:--

The Procumbent Marsh-wort (Helosciadium nodiflorum or Apium nodiflorum) is a creeping plant, abundant in ditches and other wet places, rooting at the base, with erect flowering stems that are sometimes very short, but often reach a height of three feet. The whole plant is smooth, with hollow stems; pinnate leaves with from three to nine or more pairs of ovate, bluntly-toothed leaflets; and almost sessile umbels of small, white flowers either opposite the leaves or in the angles of the upper branches. These umbels are compound, with about five or six rays, usually without primary bracts, but with several, narrow, secondary ones. The petals have their points turned inwards; and the carpels are oval, each with five narrow ribs. This plant is commonly seen growing in company with the Water-cress and the Brooklime, and blooms in July and August.

In ditches we occasionally meet with the Water Hemlock or Cowbane (Cicuta virosa)--a tall plant, from three to four feet high, bearing large, flat umbels of small, white flowers from June to August. Its stem is hollow, furrowed, and branched; and the leaves are large, twice pinnate or ternate, with lanceolate, acute leaflets, generally over an inch in length, the margins serrate or (sometimes) doubly serrate. Comparing this plant with the Common Hemlock (p. 169), we should note that the secondary bracts of the latter are three in number, almost invariably turned to the outside; and that its calyx teeth are very indistinct, while in the present species they are prominent above the ovary.

Next follow three species of Water Dropwort (genus nanthe)--smooth plants, with much-divided leaves and compound umbels of white flowers, with secondary, and sometimes also primary, narrow bracts. In all three species the central flowers of each secondary umbel are perfect and shortly stalked, while the outer ones are on longer stalks, and usually staminate. The petals are notched, with points turned inwards; and the fruits have two rather long styles, are crowned by the five minute teeth of the calyx, and their carpels have each five blunt ribs. The three species referred to are:--

1. The Common Water Dropwort (. fistulosa).--An erect plant, from two to three feet high, with a fleshy, fibrous root; creeping runners; and a thick, hollow, slightly-branched stem. Its radical leaves are bipinnate, with segments cut into three or five narrow lobes; and the stem leaves have long, hollow stalks, with a few narrow segments at the top. The umbels have from three to five rays, usually with no primary bracts, and a few, narrow secondary ones.

2. The Hemlock Water Dropwort (. crocata) is a larger plant, from two to five feet high, with a tuberous root and a thick, branched stem. Its leaves are bipinnate, with stalked, shining leaflets that are irregularly cut. The umbels are on long stalks, and have nearly twenty rays, several narrow secondary bracts, and sometimes a few primary ones. The middle flowers of each secondary umbel are perfect and almost sessile, but the outer ones are stalked and staminate.

3. The Fine-leaved Water Dropwort (. Phellandrium) grows from one to four feet high, and has an erect, creeping or floating stem with runners at the base. The upper leaves are bipinnate, with small, cut segments; and the submerged ones are deeply cut into very narrow, almost hair-like lobes. The umbels are small, on short stalks in the angles of the branches or opposite the leaves. They have about ten rays, narrow secondary bracts, but no primary ones.

All three of the above species flower from July to September.

Next follow a few composite flowers (order Compositæ), the first of which is the Marsh Thistle (Carduus palustris) that varies from two to eight feet in height, and bears dense clusters of purple (occasionally white) heads during July and August. Its stem is stiff, hollow, slightly branched, and thickly covered with very prickly wings that are continuous with the margins of leaves above them. The leaves are narrow, wavy, deeply divided into prickly lobes, with scattered hairs on both surfaces; the lower ones often seven or eight inches long; and the upper much smaller and narrower. The flower-heads are ovoid, surrounded by an involucre of many closely-overlapping bracts with prickly tips.

The two Bur Marigolds (Bidens) are more or less common in marshes and other wet places. They are both smooth plants with opposite leaves, and hemispherical heads of yellowish flowers surrounded by two or three rows of bracts, the outer of which are spreading. The receptacle is flat, with membranous scales between the florets; and the fruits are crowned by from two to five stiff, prickly bristles. The more abundant of these is the Nodding Bur Marigold (B. cernua), a stout plant, from one to two feet high, distinguished by its narrow, entire, sessile leaves, and its drooping flower-heads. The other--the Trifid Bur Marigold (B. tripartita)--has three-cleft, stalked leaves, and heads erect or only slightly drooping.

The Common Ragwort of waste places, described on p. 187, is represented in marshes and wet places by a very similar plant called the Marsh Ragwort (Senecio aquaticus), which varies from one to three feet in height, and flowers in July and August. Its stem is more slender than that of S. Jacobæa, and is usually more branched. The leaves are either deeply toothed, or pinnately cut into segments which decrease in size towards the base. The yellow flower-heads are not so densely crowded as in the Common Ragwort, and have longer stalks.

The Scrophulariaceæ contains three common plants of the Veronica genus that grow in wet places. All three are similar in that they have opposite leaves; a corolla with a short tube, and four spreading limbs, of which the lowest is narrowest; two stamens; and a capsular fruit, flattened at right angles to its partition, opening by two valves, and containing a few seeds.

One of these is the Marsh Speedwell (Veronica scutellata), abundant in the marshes and ditches of most parts of Britain. It has a weak, straggling stem, from four to eight inches high, with creeping runners at the base; narrow, smooth, sessile leaves, either uncut or only slightly toothed; and slender racemes of pale pink or white flowers on axillary peduncles arranged alternately, there being only one raceme at each node.

The second is the Water Speedwell (V. Anagallis), a smooth plant, varying from six inches to two feet high, abundant in marshes and ditches, bearing small lilac or white flowers in July and August. Its stem is stout, succulent, hollow, erect, and slightly branched; the leaves narrow, acute, toothed, sessile, sometimes clasping the stem; and the racemes axillary and opposite. The flowers are only a fifth of an inch across.

The third is the Brooklime (V. Beccabunga), a very abundant plant commonly seen growing in ditches in company with the Water Cress and the Marsh-wort. It is a smooth plant, with a stem from one to two feet long, procumbent at the base and rooting at the joints; erect, succulent flowering branches; thick, elliptical, blunt, slightly-toothed leaves on short stalks; and opposite, axillary racemes of blue (occasionally pink) flowers about a third of an inch across.

Two of the Figworts, belonging to the same order (Scrophulariaceæ), are abundant in wet places all over Britain. They are both tall erect plants, with opposite leaves, and peculiar greenish brown or dull purple flowers. In both the corolla is almost spherical and shortly lipped. Two of the five lobes form the upper lip; two are at the sides; and the other, forming the lower lip, is turned down. There are five stamens, four of which are fertile and turned down, while the fifth is barren and scale-like, under the upper lip of the corolla.

One species--the Water Figwort (Scrophularia aquatica)--grows in marshes and on the banks of ditches and streams. It has a stout, angular stem, the angles of which are drawn out into narrow wings; smooth, opposite, blunt leaves, cordate at the base, with crenate or toothed margins; and long, narrow panicles of flowers with blunt bracts. The five lobes of the calyx are fringed with a conspicuous, transparent, membranous border.

The other is the Knotted Figwort (S. nodosa), which is much like the last, but emits a disagreeable odour, and may be further distinguished by the little green, fleshy knots of its rhizome. Its stem is sharply four-angled, but not winged; its leaves are acute, and doubly toothed; and the panicle has small, narrow, sharp bracts.

Passing now to the order Labiatæ, we come first to the Gipsy-wort (Lycopus europæus), an erect, branched, slightly hairy plant, from one to three feet high, bearing dense whorls of small, white, sessile flowers from June to September. The calyx has five equal teeth with stiff points; and the corolla, which is only slightly longer than the calyx, has four nearly equal lobes. This plant is abundant in most parts of Britain, and is generally seen on the banks of ditches.

In the same order we have the Mints (genus Mentha)--strongly-scented plants with creeping rootstocks and runners; and small flowers in dense, axillary whorls, or in terminal spikes or clusters. In all the calyx has five equal teeth; and the corolla is bell-shaped, with a short tube, and four lobes of which the upper is broader. There are four erect, equal stamens; and the fruit consists of four small, smooth nuts. Three species, more or less abundant, occur in marshy or other wet places. They are:--

1. The Round-leaved Mint (Mentha rotundifolia).--A moderately common, erect, hairy plant, from one to three feet high, with a powerful but hardly agreeable odour. Its stem is green, hairy, and branched; and the leaves are sessile, broadly ovate or round, blunt, wrinkled, green above, and whitish and shaggy beneath. The flowers are small, lilac (occasionally white), in dense, cylindrical, leafy spikes from one to two inches long. The bracts are rather narrow and sharply pointed, and the corolla is hairy. The time of flowering is August and September.

2. The Water Mint (M. aquatica).--An abundant marsh plant, from one to three feet high, flowering from July to September, possessing a strong, pleasant odour. Its stem is much branched, generally clothed with soft hairs; and its leaves are stalked, ovate, serrate, the upper ones passing into bracts which are shorter than the flowers. The latter are lilac, and form dense, terminal, oblong or globular clusters, with, frequently, two or three dense, axillary whorls beneath. The calyx is tubular, about an eighth of an inch long, with very sharp teeth.

3. The Marsh Whorled Mint (M. sativa).--A very similar plant, common in wet places, flowering during July and August. It grows from two to five feet high; and its elliptical, toothed leaves are hairy on both sides. The flowers are lilac, in dense, axillary whorls, without any terminal cluster.

There is yet another marsh plant of the Labiatæ to be considered, and that is the Marsh Woundwort (Stachys palustris), which is very much like the Hedge Woundwort described on p. 199. It has a stout, hollow, hairy stem, from one to three feet high; and narrow, coarsely-toothed leaves, from two to four inches long, the upper ones sessile and the lower shortly stalked. The flowers are pale purple or dull, light red, arranged in whorls of from six to ten in the axils of the upper leaves. The calyx is bell-shaped, with ten ribs and five long, acute teeth; and the lower lip of the corolla has its side lobes turned back.

We now reach the interesting Myosotis genus of the Boraginaceæ, containing the favourite Forget-me-not and the similar Scorpion-grasses. They are all rather low and weak plants, with small, sessile, narrow leaves; and small flowers in one-sided, curved racemes without bracts. The calyx is cleft into five; and the corolla has a short tube, partially closed by five little scales, and five spreading or concave lobes. The stamens are enclosed in the tubes of the flower. Three species are common in wet places. They are--

1. The Forget-me-not (Myosotis palustris).--An abundant plant, growing to a foot or more in height, and bearing, from June to August, bright blue flowers, nearly half an inch across, with a yellow centre. It has a creeping rootstock, with runners, and rather weak ascending stems clothed with spreading hairs. The leaves are blunt, and often covered with hairs that lie close against the surface. The calyx is divided to about a third of its length into short, triangular teeth, and is covered with closely-pressed hairs.

2. The Creeping Water Scorpion-grass (M. repens).--A very similar plant, sometimes regarded as a variety of the last. Its stock emits leafy runners above the ground, and the stem is more hairy. The flowers, too, are of about the same size, but of a sky-blue colour, and their stalks are longer, bending downwards when in fruit. The calyx is divided to about the middle into narrow teeth.

3. The Tufted Water Scorpion-grass (M. cæspitosa).--Also a similar plant, often regarded as a variety of M. palustris; but its flowers are only about half the size, of a sky-blue colour, with narrow calyx teeth almost as long as the corolla. It is of a paler green colour, and the stems are tufted by a free branching at the base.