Part 34

3. The Common St. John's-Wort (H. perforatum).--A very common plant in woods and thickets, growing from one to two feet high, and flowering from July to September. It has short, underground stems, or barren shoots that lie on the ground and root at the nodes, in addition to the erect, flowering stems, which are either round or two-edged, and branched towards the top. The leaves are half an inch long, with opaque veins, many transparent dots, and sometimes a few black dots on the under side. The yellow flowers form a terminal corymb. Their sepals are narrow, about half the length of the petals. The stamens are in three sets, united at the base; and both petals and anthers are marked with black dots.

4. The Hairy St. John's-Wort (H. hirsutum).--A stiff, erect plant, from one to three feet high, common in the woods and thickets of most parts of Britain, flowering in July and August. The stem is round, and clothed with soft hairs. The leaves are ovate, oblong or elliptical, tapering at the base into a short stalk, about an inch long, with many transparent dots, and downy along the veins on the under side. The sepals are narrow, acute, about half the length of the yellow petals, and fringed with stalked glands. The stamens are in three sets.

The Wood Crane's-bill (Geranium sylvaticum--order Geraniaceæ), one of the most handsome of our Wild Geraniums, is not found in the South, but is moderately common in parts of North Britain, including North Ireland. Its stem is erect, from one to two feet high, and branched towards the top. In general form the leaves are heart-shaped or shield-shaped, but they are very deeply divided into five or seven radiating, cut, and toothed lobes. The lower ones are on long stalks; but the upper are shortly stalked or sessile, and less divided. The flowers are of a bluish-purple or rose colour, about an inch in diameter, arranged in a loose panicle with two flowers on each pedicel. The five sepals are about half the length of the petals, and terminate in a very fine point; and the petals are obovate and slightly notched. The plant flowers during June and July.

Passing now to the order Leguminosæ we note first the Dyer's Green-weed (Genista tinctoria)--a shrubby plant, common in the thickets and bushy places of South Britain, flowering from July to September. The stem is woody and stiff, but green; its base rests on the ground, but it sends up erect flowering branches from one to two feet high. The yellow flowers are arranged in terminal racemes, each flower having a lanceolate bract at the base of its short stalk, and very small bracts at the base of the calyx. The calyx has five teeth, the three lower ones much narrower than the other two, all terminating in a sharp point; the corolla is much longer than the calyx, with an oblong standard or upper petal; the stamens are all united by their filaments, forming a complete sheath round the ovary; and the pods are smooth, about an inch long, and compressed.

In the thickets of most parts of Britain, but more especially those of the eastern counties, we may often meet with the Sweet Milk Vetch (Astragalus glycyphyllos) of the same order. It is a prostrate plant, with pale yellow or cream-coloured flowers that bloom from June to August. The flowers are about half an inch long, in short, dense, shortly-stalked racemes. The calyx has five teeth; the upper stamen is free from the other nine, which form a divided sheath round the ovary; and the pod is smooth, round, curved, over an inch long, and divided by a double membrane into two cells, each of which contains about seven seeds.

In the same order are two species of Everlasting Pea (Lathyrus), both of which grow in thickets and other bushy places. One is the Tuberous Everlasting Pea or Tuberous Bitter Vetch (L. macrorrhizus), an erect plant, from six inches to a foot in height, flowering from May to July. Its rootstock has small tubers, and the stem is winged. The leaves are pinnate, with from two to four pairs of narrow leaflets and half arrow-shaped stipules; they have no tendrils, but the leafstalk terminates in a fine point. The flowers are of a red-purple colour, changing to greenish blue as they fade; and are in loose racemes of from two to four.

The other is the Narrow-leaved Everlasting Pea (L. sylvestris), a straggling plant, from two to six feet long, flowering from June to August. It is not so common as the last, but may be found in similar situations. Its stem has very narrow wings; and the leaves have very narrow leaflets, flattened stalks, branched tendrils, and half arrow-shaped stipules. The flowers are rather large, of a pale purple colour, with a greenish keel, and a green spot on the large upper petal. They are arranged in loose racemes.

The Wild Raspberry (Rubus Idæus--order Rosaceæ) is to be found in the woods and thickets of most parts of Britain. It may be easily distinguished from other species of its genus by the following description:--Rootstock creeping, with many suckers. Stems round, erect, with a soft down and numerous weak prickles. Leaves pinnate, with three or five ovate, pointed, toothed leaflets, pale green above, and white and hoary beneath. Stipules small, very narrow and pointed, usually attached part way up to the leafstalk. Flowers white, in long, terminal, drooping panicles. Calyx five-lobed; petals five, short and narrow; stamens numerous; and fruit consisting of a globular cluster of red or yellow, hoary, one-seeded, succulent carpels which usually separate from the conical receptacle when ripe. The bush grows from three to five feet high, and flowers from June to August.

Two species of Willow Herb (order Onagraceæ) grow in copses and thickets, and are easily recognised by their rose-coloured flowers with very long, inferior ovaries. One is the beautiful French Willow or Rose Bay Willow Herb (Epilobium angustifolium), an erect plant, varying from two to six feet in height, widely distributed, though not very common, flowering during July and August. Its leaves are alternate, narrow-elliptical, entire or with very small teeth, and very shortly stalked. The flowers are about an inch in diameter, numerous, forming a very long, loose, terminal, tapering raceme, with a narrow bract at the base of each pedicel. The calyx is tubular, four-cleft, attached to the top of the long ovary; the corolla consists of four entire, nearly equal, spreading petals; the stamens, eight in number, all bend downwards; and the stigma is deeply divided into four lobes, on a long style which also bends downward. The fruit is a four-celled capsule, two or three inches long, which splits when ripe, its valves curling downwards and exposing numerous minute seeds, each of which has a silky tuft of fine hairs that enables it to be dispersed by the wind. The plant is most frequently seen in damp copses, and among the undergrowth of damp woods.

The second species is the Pale Smooth-leaved Willow Herb (E. roseum), an erect plant, seldom more than two feet high, found principally in the damp copses of the southern counties, flowering in July and August. Its stem is four-angled, two opposite angles being much more prominent than the other two; and its leaves are opposite, with longer stalks, lanceolate or elliptical, pointed, toothed, smooth, usually about two inches long. The flowers are not nearly so numerous as those of the last species, are only a little over a third of an inch in diameter, and in a short, leafy panicle, drooping while in the bud. The calyx is deeply divided into four sepals about a sixth of an inch long; the corolla consists of four notched petals, a little longer than the sepals; the stamens, ovary, fruit, and seeds correspond in number and character with those of the last species; but the stigma is either entire or divided into four very short lobes.

In the same order we have the Enchanter's Nightshade (Circæa lutetiana), distinguished at once from the Willow Herbs by having only two sepals, two petals, and two stamens. It is an erect, hairy plant, from one to two feet high, flowering from June to August. Its stem is slender; and the leaves are opposite, long-stalked, ovate and coarsely toothed. The flowers are very small, white, in terminal, leafless racemes, with deeply-notched petals, and pink stamens. The fruit is a little two-lobed capsule with stiff, hooked hairs.

The Cornel or Dogwood (Cornus sanguinea), of the order Cornaceæ, is a common shrub in woods and thickets, and is often employed in the making of hedgerows. It grows from five to eight feet high, and flowers during June and July. Its leaves are covered, when young, with fine, silky hairs that lie close on the surface, but these almost entirely disappear later; and towards the end of the summer the leaves assume a deep crimson or purple colour. The flowers are very abundant, of a yellowish white colour, and are arranged in dense cymes, about two inches across, without bracts. The four-toothed calyx and the peduncle are both clothed with a mealy down; and the four petals, about a quarter of an inch long, are narrow and pointed. The fruit is a purple-black, globular, berry-like drupe, containing a stone with one or two seeds.

In very dense woods, where the light is so much reduced that but few flowers will grow, we may generally find the Wood Sanicle (Sanicula europæa), a smooth umbelliferous plant with a short, hard rootstock, and a simple stem from one to two feet high. The leaves, which are all radical, are on long stalks, and are palmately divided into three or five shining lobes that are themselves cut and sharply toothed. The flowers are sessile, in little rounded heads; the whole inflorescence forming an irregular umbel or a loose panicle. They are very minute, of a pinkish white colour; and the outer ones of each head usually have no pistil. They bloom during June and July, and are followed later by little prickly fruits about a sixth of an inch long.

In damp woods we commonly meet with the tall, stout, branching Angelica (Angelica sylvestris) of the same order (Umbelliferæ), with a thick, furrowed stem, two to four feet high, downy above, and usually more or less shaded with purple. Its lower leaves are very large, with stalked, ovate leaflets, from one to two inches long, often three-lobed, and always sharply toothed. The upper leaves are much smaller, with fewer leaflets, and often consist only of a broad sheath with a few small leaflets at its summit. The flowers are white, generally tinged with pink, and form a large terminal umbel of from sixteen to forty rays, with two or three narrow primary bracts, and several fine secondary ones. They bloom during July and August, and are succeeded by flattened fruits with three ribs on the back of each of the two carpels. The carpels are also broadly winged; and, as the wings do not adhere, each fruit is surrounded by a double wing.

The order Caprifoliaceæ includes the Common Elder (Sambucus nigra), the white or cream-coloured flowers of which are so conspicuous in our woods and hedgerows in June. This tree grows to a height of fifteen or twenty feet, and its young branches are remarkable for the large quantity of pith they contain. The general form of the leaves and the arrangement of the flowers are seen in our illustration. Each flower has a calyx with five small teeth; a corolla with a short tube and five spreading limbs; five stamens attached to the base of the corolla; and an inferior ovary. The fruit is a black, berry-like drupe containing (usually) eight little, seedlike stones.

The Guelder Rose or Water Elder (Viburnum Opulus), of the same order, is a flowering shrub, usually six or eight feet high, moderately common in moist woods and copses, especially in the South, bearing showy cymes of white blossoms in June and July. The cymes are flat-topped, consisting of numerous flowers, the outer of which are much larger, often nearly an inch in diameter, but without stamens or styles, while the others are perfect, with five stamens and three sessile styles. The fruit is a blackish-red, almost globular, slightly-flattened berry, containing a single seed. The cultivated variety of this shrub, known as the Snowball Tree, has large, globular cymes of flowers, all of which are large and barren.

The Great Valerian or All-heal (Valeriana officinalis--order Valerianaceæ) is moderately common in moist woods, and is rather widely distributed. It is an erect plant, from two to four feet high, flowering from June to August. There seems to be two distinct varieties of this plant, one with from four to six pairs of leaflets, and the other with from six to ten pairs, in addition to the terminal leaflet in each case. The flowers are small, flesh-coloured or nearly white, in terminal and axillary corymbs. The little inferior ovary is surmounted by a calyx which is compactly rolled in at first, but which expands into a spreading, feathery pappus as the fruit ripens. The corolla is tubular, with five short, equal, spreading lobes. It is not spurred as in the case of the Red Valerian (p. 302), but the base of the tube is pouched on one side. This plant is shown on Plate II, Fig. 1.

We have now to note some composite flowers (order Compositæ) of wooded and shaded ground. Of these we will first take the Blunt-leaved or Succory-leaved Hawk's-beard (Crepis succisæfolia or C. hieracoides), which is moderately common in the woods of North England and Scotland, but does not occur in the South. It is an erect plant, varying from one to three feet in height, smooth or slightly hairy, flowering during July and August. The fruits (achenes) are marked by many fine, longitudinal ridges, and are surmounted by a dense pappus of soft, white hairs which are a little longer than the fruits themselves. This flower is represented in Plate II, Fig. 3.

In the woods and thickets of nearly all parts of Britain we may see the Saw-wort (Serratula tinctoria), a stiff, erect, smooth plant, from one to three feet high, flowering in August and September. The flower-heads are purple or crimson, forming a loose, terminal corymb; and the florets, all of which are tubular, are imperfect, the males and females being generally on different plants. The involucre is oblong in form, more than half an inch long, consisting of many pointed, closely-placed bracts, of which the inner are usually tipped with red; and that of the male heads is somewhat broader than the involucre of the females. The pappus consists of a tuft of simple hairs, most of which are longer than the achene.

The Golden Rod (Solidago Virga-aurea) is another abundant flower, found in dry woods and thickets in all parts. It is a tufted plant, with stiff, erect, angular, slightly-branched stems, varying from six inches to two feet in height; and narrow-elliptical leaves, entire or slightly toothed, the lower ones stalked. The flowers are very numerous, of a bright golden yellow-colour, forming a dense, terminal panicle. The heads are not large, and each consists of about twenty tubular disc-florets; half the number of strap-shaped ray-florets; and an involucre of many overlapping bracts. The pappus consists of many simple hairs. This species flowers from July to September.

Two species of Leopard's Bane (Doronicum) are occasionally to be seen in damp woods and thickets, especially near villages. They are not indigenous, only occurring as escapes from gardens, but they have now become well established as wild flowers in many parts of Britain. Both are tall, erect plants, from two to three feet high, with large yellow heads surrounded by two or three rows of narrow, acute bracts. Except in colour the heads much resemble the Ox-eye Daisy. In both species the achenes of the ray have no pappus, but those of the disc have a pappus of stiff hairs in several rows. They flower from May to July.

The Great Leopard's Bane (D. Pardalianches) has a creeping rootstock and a hollow stem. Its radical leaves are broadly heart-shaped, slightly toothed, on long stalks; and the stem leaves are narrower, entire or toothed, the upper ones small, sessile, embracing the stem; and the lower ones stalked, with a broad expansion at the base of the stalk which clasps the stem. The heads are usually three or four in number, on long leafless peduncles.

The other species, the Plantain-leaved Leopard's Bane (D. plantagineum), has, as its name denotes, leaves similar to those of the Plantain. It usually has solitary flower-heads, and is represented on Plate I.

Passing now to the favourite Bell-flowers (Order Campanulaceæ), we have to notice four species that are to be found in woods and other shady spots during the summer months. The features common to the four species are:--Leaves alternate. Calyx adhering to the ovary, with a border of five lobes or teeth. Corolla bell-shaped, with five lobes. Stamens five, attached to the corolla by the broad bases of the filaments. Ovary inferior, ripening to a capsule that opens by longitudinal clefts. The species referred to are:

1. The Giant Bell-flower (Campanula latifolia). A stout plant, from three to five feet high; with an unbranched, leafy stem; and a leafy raceme of large, deep blue or white flowers that bloom in July and August. Its leaves are large, ovate to lanceolate, acute, doubly serrate, the lower ones stalked and the upper sessile. Each axillary peduncle bears only one flower, the calyx of which has long, narrow segments, and the corolla is hairy within. The capsule is short, opening by slits near the base. This flower is found principally in the North.

2. The Creeping Bell-flower (C. Rapunculoides).--A downy plant, with a creeping rootstock; an erect, simple or slightly-branched stem from one to two feet high; and a one-sided raceme of drooping, deep blue flowers that appear in July and August. The leaves are rough and doubly toothed, the lower ones stalked and heart-shaped, and the upper narrow and sessile. The segments of the calyx are long and narrow, and the capsule is globular, opening by small slits near the base. This species is widely distributed, but not very common.

3. The Nettle-leaved Bell-flower (C. Trachelium).--A very rough plant, with an angled stem, from one to three feet high, bearing a leafy raceme of large blue flowers from July to October. Its leaves are much like those of the Stinging Nettle, being very rough, bristly, and coarsely toothed. The segments of the calyx are rather broad, and very rough with stiff hairs. This species is very abundant in some localities, and is widely distributed. (See Plate II, Fig. 4.)

4. The Ivy-leaved Bell-flower (C. hederacea).--A pretty little creeping plant that grows in moist woods, flowering during July and August. It is very widely distributed, and is a common flower in many parts of Great Britain, more especially in the southern counties. Its prostrate stem is very slender; and the leaves are small, stalked, very broad, and palmately divided into angular lobes. The flowers are of a pale blue colour, solitary on long, threadlike peduncles; and the capsule is globular, opening by three valves at the top.