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The Zoology of the Voyage of H.m.s. Beagle, [vol. 4 of 5] · Leonard Jenyns — chapter 14 of 60 · ~2,039 words · public domain

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B. 7; D. 10—1/25; A. 1/8; C. 17; P. 21; V. 1/5. Length 9 inches 6 lines.

COLOUR.—“Body mottled with silver and green: dorsal and caudal fins lead-colour.”—D. In spirits, the colour appears dusky brown, with darker mottlings and silvery reflections; paler beneath. The fins are dusky, but the basal half of the dorsal is darker than the upper. The pectorals are darker than the other fins, especially the inside; on the left pectoral, the dark colour is restricted to three broad transverse fasciæ. There are also on the pectorals and anal, and on most of the scales on the body, small blackish dots, as mentioned in the ‘Histoire des Poissons.’

A second specimen, smaller than the above, has the back rather more arched, the greatest depth being only five times in the length. The outer row of teeth in the upper jaw is not quite so conspicuous, the teeth being smaller and closer-set, and consequently more numerous. The soft dorsal and anal have fewer rays.

D. 10–1/22; A. 1/7; &c. Length 7 inches 3 lines.

In all other respects similar to the specimen first described.

Habitat, Bahia Blanca, and Maldonado.

As Cuvier and Valenciennes have mentioned individuals of this species, which varied in the number of rays in the soft dorsal from twenty-two to twenty-four, I cannot but consider the two above described as specifically the same, though in the first these rays amount to as many as twenty-five. This, which is the larger specimen, was taken by Mr. Darwin at Bahia Blanca, where it is said to have been common. The other was obtained at Maldonado.

2. UMBRINA OPHICEPHALA. Jen.

U. elongata; rostro obtusissimo, tumido, haud ultrà fauces producto, margine inferiore quadrilobato, lobis intermediis rotundatis; fossulá longitudinali inter nares, profundè exaratâ; poris quatuor infrà symphysin; dentibus velutinis, serie externâ in maxillâ superiore aculeiformi; preoperculo obsoletè denticulato; operculo mucronibus duobus parvis instructo; spinis dorsalibus tertiâ et quartâ longissimis, corporis altitudinem æquantibus; spinâ anali gracili, radiis articulatis dimidio breviori.

D. 12—1/22; A. 1/9; C. 17; P. 20; V. 1/5. LONG. unc. 6. lin. 5.

FORM.—Very much elongated; the greatest depth just one-sixth of the entire length; the head one-fifth. Dorsal line nearly straight. Profile falling very slightly till it reaches the nostrils, when it suddenly becomes vertical. Snout in consequence short, and very blunt, and not projecting beyond the jaws; with a deep broad channel down the middle, extending from between the nostrils to near the mouth: on each side of this channel, the snout is very protuberant. The lower margin of the snout is divided into four lobes, the central pair of which are rounded: above each of the exterior lobes is one large pore, and an odd one in the middle. There are also four pores beneath the symphysis, and a short barbule, as in the last species. The eye has a diameter about one-fifth the length of the head, and is distant one diameter from the end of the snout. The nostrils, which are immediately in advance of the eye, consist of two round apertures, one before the other, the posterior one double the size of the anterior. Upper jaw a very little longer than the lower. A band of velutine teeth in each jaw; with an outer row above of moderately strong card, rather curving inwards and backwards, and closer-set than those of the U. arenata, amounting to twenty-eight or thirty in number: there are also some smaller card teeth behind this outer row passing insensibly into the velutine. Preopercle very obsoletely denticulated. Opercle with two flat points not very obvious.

First dorsal triangular, and moderately high in the point; third and fourth spines longest, about equalling the depth of the body; first spine very small: all the spines rather slender. Second dorsal about half as long again as the first, but the rays are too much broken to judge of their relative lengths. Anal spine very slender, and about half the length of the soft rays. The caudal is injured, but appears to have been of nearly the same form as in the U. arenata. The pectorals are about three-fourths the length of the head, but the ends of the rays are worn. The ventrals are of the same length as the pectorals in their present state: they are placed rather backwarder than in the U. arenata, being attached beneath the first third of the pectorals: there is a pointed scale in their axilla, of about the same relative size as in that species. The scales on the body are rather smaller, ciliated on their free edges, with a fan of eleven or twelve striæ behind. There are rows of small scales on the caudal, but none apparent on the other fins.

COLOUR.—Mr. Darwin did not notice the colours of this species in its recent state. In spirits, it appears of a nearly uniform dusky brown, but paler on the abdomen, with traces of silvery reflections about the head. The fins are dark, but the anal paler at the base than at the tips of the rays.

Habitat, Coquimbo, Chile.

This species may be at once distinguished from all those described in the ‘Histoire des Poissons,’ by its very elongated form. The head also has a peculiar character about it, and is not unlike that of some serpents. It appears to be the first species of this genus brought from the Pacific, the other foreign ones being all found either in the Indian seas, or on the Atlantic side of America. There are two specimens in the collection, exactly similar, and both obtained by Mr. Darwin at Coquimbo. They are, however, both in very bad condition; so much so, indeed, that I should have hesitated about describing them as new, had they not presented several obvious peculiarities.

GENUS—PRIONODES. Jen.

Serrani formam quam maximè gerens. Pinna dorsalis unica, per totam longitudinem subæqualis. Membrana branchialis septem-radiata. Nec fovea, nec pori, infrà symphysin. Dentes maxillares velutini, serie externâ cæteris fortiori, paucis, hic illic sparsis, subcaninis; palatini nulli. Preoperculum denticulatum. Operculum mucronibus tribus posticè armatum. Spina analis secunda fortis. Squamæ corporis ciliatæ; minutissimæ inter radios pinnarum verticalium, in seriebus dispositæ.

Fish. Pl. 9.

W. Hawkins del^t. ]

Fig. 1. Prionodes fasciatus. 2. Stegastes imbricatus. Nat. Size.

I am called upon either to establish this new genus among the Sciænidæ, or to break down one of the essential distinctions set by Cuvier between this family and the Percidæ. The form is so completely that of a Serranus,—which it resembles especially in its dorsal fin, head, maxillary teeth, form and armature of the pieces of the gill-cover, and in the arrangement of the scales on the body,—that at first sight no one would hesitate to refer it to that group; but the vomer and palatines are without teeth. In this respect, indeed, I consider it an important discovery; as it affords another striking instance of the uncertainty of this character, in cases in which others, which have been generally made subordinate to it, remain constant. It is probable that the time will come, when it will be found necessary to revise some portion of the Percidæ and Sciænidæ with reference to a more correct valuation of this character. For the present, however, I refrain from interfering with the Cuvierian arrangement; and the only alternative is to consider this as a new form among the Sciænidæ, where it must be placed along with those genera possessing one dorsal fin, and having seven rays in the branchiostegous membrane. Such are Hæmulon, Pristipoma, and Diagramma; from all which, however, it is at once distinguished by the absence of pores at the symphysis and on the lower jaw, and by the much more developed spines on the opercle, and from Pristipoma by its having, further, scales on the vertical fins. On the whole, it seems to approach nearest to Hæmulon; but the crown and snout are more free from scales than in that genus, and the scales on the body are not set in oblique rows, as is the case in so many of the true Sciænidæ. The head also has no cavernous appearance about it. This new form is from the Galapagos Archipelago.

PRIONODES FASCIATUS. Jen.

PLATE IX. Fig. 1.

P. pallidè flavescenti-fuscus, fasciis transversis plurimis suprà rubescenti-nigris infrà miniatis; pinnis verticalibus maculis parvis ocellatis: vertice, rostro, et maxillis, nudis; preoperculo margine adscendenti denticulato, basali lævi; operculo mucrone intermedio forti; spinis dorsalibus ad apices laciniis investitis; pinná caudali subæquali.

B. 7; D. 10/12; A. 3/7; C. 17; P. 18; V. 1/5. LONG. unc. 7. lin. 3.

FORM.—Oval, compressed; the back not much arched, forming one continuous curve with the profile, which falls gently from the nape; ventral line less convex than the dorsal. Greatest depth equalling one-fourth of the entire length; head about one-third; thickness rather less than two-thirds of the depth. Mouth rather wide, with the lower jaw longest. The maxillary dilates at its posterior extremity; it reaches to nearly beneath the middle of the orbit, and does not retire beneath the suborbital. In each jaw a band of velutine teeth; above there is an outer row of longer ones in card, and one or two in front on each side still longer resembling small canines; in the lower jaw there are also a few longer ones, of the same character as these last, interspersed at intervals. Tongue free at its extremity, and, as well as the vomer and palatines, without teeth. Eyes rather high in the cheek; their diameter about one-sixth that of the head. The nostrils consist of two small round orifices a little in advance of the eyes, the anterior one covered by a membranous flap. Margin of the suborbital entire. Preopercle finely denticulated on its ascending margin, which is vertical and slightly convex; but the denticulations almost disappear at the angle, and are not visible at all on the basal margin. Opercle triangular, with three flat spines, the middle one longest, beyond which the membrane projects in the form of an angular process to the distance of three lines. Small scales on the cheeks and preopercle; but none on the crown, snout, first suborbital, maxillary, or lower jaw; scales on the opercle larger, equalling those of the body in size. Gill-opening large, with the branchial membrane deeply notched in the middle.

Lateral line following the curvature of the back at one-fourth of the depth. Scales on the body moderately large: one taken from the middle of the side above the lateral line is of a somewhat oblong form, with the free edge rounded and finely ciliated; the basal portion with fourteen slightly converging striæ, which form at the hinder margin as many, but not very distinct, crenations.

The dorsal commences above the terminating lobe of the opercle, and reaches to within a short space of the caudal: height of the spinous portion, which, with the exception of the first two spines, is nearly even throughout, about one-third of the depth; soft portion rather higher, with the last two rays but one longest, and forming a point backwards; all the soft rays branched. Anal commencing in a line with the soft portion of the dorsal, and terminating a little before that fin; three spines, the second one-third longer than the first, and a little longer than the third, and much the strongest of all; the soft portion of the anal is similar to that of the dorsal, and terminates in like manner in a point behind. Space between the anal and caudal a little less than one-sixth of the entire length. The caudal appears to have been nearly even, or perhaps slightly rounded, but the rays are worn. Rows of very minute scales, not very obvious, between the rays of all the vertical fins. Pectorals slightly rounded; more than half the length of the head; all the rays with the exception of the first two and the last, branched. Ventrals attached beneath, or perhaps a very little in advance of the pectorals; pointed, with the second soft ray longest. No lengthened scale or process of any kind in the axilla of either ventrals or pectorals; neither are the former fastened to the abdomen by a membrane half their own length, as is the case in many of the Serrani.

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