Part 2

PREFACE.

His Majesty’s ship, Beagle, under the command of Captain FitzRoy, was commissioned in July, 1831, for the purpose of surveying the southern parts of America, and afterwards of circumnavigating the world. In consequence of Captain FitzRoy having expressed a desire that some scientific person should be on board, and having offered to give up part of his own accommodations, I volunteered my services; and through the kindness of the hydrographer, Captain Beaufort, my appointment received the sanction of the Admiralty. I must here, as on all other occasions, take the opportunity of publicly acknowledging with gratitude, the obligation under which I lie to Captain FitzRoy, and to all the Officers on board the Beagle, for their constant assistance in my scientific pursuits, and for their uniform kindness to me throughout the voyage. On my return (October, 1836) to England, I found myself in possession of a large collection of specimens in various branches of natural history; but from the great expense necessary to secure their publication, I was without the means of rendering them generally serviceable.

The Presidents of the Linnean, Zoological, and Geological Societies, having given me their opinion respecting the utility to be derived from publishing these materials, I addressed a letter to the Right Honourable the Chancellor of the Exchequer (T. Spring Rice, Esq.) informing him of the circumstances under which I hoped that I might venture to solicit the aid of Government. In reply, I received a communication (as below) announcing to me that the Lords of the Treasury, from their readiness to promote Science, were willing, under certain conditions, to give me the most liberal assistance.

“Treasury Chambers, August 31, 1837.

“SIR,

“It having been represented to the Lords Commissioners of Her Majesty’s Treasury, from various quarters, that great advantage would be derived to the Science of Natural History, if arrangements could be made for enabling you to publish, in a convenient form, and at a cheap rate, the result of your labours in that branch of science, my Lords will feel themselves justified in giving their sanction to the application of a sum, not exceeding in the whole one thousand pounds, in aid of such a publication; upon the clear and distinct understanding that the Work should be published, and the plates engraved, in such a manner as to be most advantageous to the Public at large, upon a plan of arrangement to be previously submitted to, and sanctioned by the Board, after consultation with those persons, who, from their attainments in this branch of science, are the most capable of advising their Lordships thereupon; and that the payments on account of the said sum of one thousand pounds are to be made to you from time to time, on a certificate that such progress has been made in the engravings, in accordance with the plan previously approved of, as to justify the issue then applied for. My Lords have therefore directed me to communicate to you the views they entertain upon this subject; and to apprize you that they will be prepared to act in conformity with their arrangement, upon learning from you that you are ready to proceed with the Work upon the principles above laid down, and upon receiving from you a statement of the manner in which you think the Work should be published, and the plates engraved, so as most effectually to accomplish the object my Lords have in view, in sanctioning the payment from the Public Funds, in aid of the expenses of the Work in question.

“I remain, “Sir, Your Obedient Servant, “A. Y. SPEARMAN.”

The object of the present Work is to give descriptions and figures of undescribed and imperfectly known animals, both fossil and recent, together with some account, in the one case, of their geological position, and in the other of their habits and ranges. As I do not possess the knowledge requisite for such an undertaking, and as I am, moreover, particularly engaged in preparing an account of the geological observations, made during the voyage, several gentlemen have most kindly undertaken different portions of the Work. Besides the very great advantage insured in thus enlisting the attainments of these Naturalists in the several departments of science, to which they have paid most attention, a great delay is avoided by adopting this method of publication, which must otherwise have been incurred before the materials could have been made known.

An Account of the Voyage, drawn up by Captain FitzRoy, (and to which I have added a volume) being on the point of publication, I shall not in this Work enter on any minute details respecting the countries which were visited, but shall merely give a sketch of the geology in the introduction to the part containing Fossil Mammalia, and a brief geographical notice in that attached to the account of existing animals. At the conclusion of this Work, I shall endeavour to place together the leading results in the natural history of the different countries, from which the collections were procured. I may here state that Mr. Owen has undertaken the description of the Fossil Mammalia; Mr. Waterhouse, the Recent Mammalia; Mr. Gould, the Birds; Mr. Bell, the Reptiles; and the Rev. L. Jenyns, the Fish. Whatever assistance I may obtain in the invertebrate classes, will be noticed in their respective places. The specimens have been presented to the various public museums, in which it was thought they would be of most general service: mention will be made in each part where the objects described have been deposited.

FOSSIL MAMMALIA,

Described by

RICHARD OWEN, ESQ. F.R.S. F.G.S. F.L.S.

PROFESSOR OF ANATOMY AND PHYSIOLOGY TO THE ROYAL COLLEGE OF SURGEONS IN LONDON; CORRESPONDING MEMBER OF THE ROYAL ACADEMY OF SCIENCES OF BERLIN; OF THE ROYAL ACADEMY OF MEDICINE, AND PHILOMATHIC SOCIETY OF PARIS; OF THE ACADEMY OF SCIENCES OF PHILADELPHIA, MOSCOW, ERLANGEN.

WITH

A GEOLOGICAL INTRODUCTION,

BY CHARLES DARWIN, ESQ. M.A. F.G.S. &c. &c.

CORRESPONDING MEMBER OF THE ZOOLOGICAL SOCIETY.

GEOLOGICAL INTRODUCTION.

BY MR. DARWIN.

Mr. Owen having undertaken the description of the fossil remains of the Mammalia, which were collected during the voyage of the Beagle, and which are now deposited in the Museum of the College of Surgeons in London, it remains for me briefly to state the circumstances under which they were discovered. As it would require a lengthened discussion to enter fully on the geological history of the deposits in which these remains have been preserved, and as this will be the subject of a separate work, I shall here only give sufficient details, for the reader to form some general idea of the epoch, at which these animals lived,—of their relative antiquity one to the other,—and of the circumstances under which their skeletons were embedded. All the remains were found between latitudes 31° and 50° on the eastern side of South America. The localities may conveniently be classed under three divisions, namely—the Provinces bordering the Plata; Bahia Blanca situated near the confines of Northern Patagonia; and Southern Patagonia.

The first division includes an enormous area, abounding with the remains of large animals. To the eastward and southward of the great streams, which unite to form the estuary of the Plata, those almost boundless plains extend, which are known by the name of the Pampas. Their physical constitution does not vary over a wide extent;—the traveller may pass for many hundred miles on a level surface, without meeting with a single pebble, or discovering any change in the nature of the soil. The formation consists of a reddish argillaceous earth, generally containing irregular concretions of a pale brown, indurated marl. This stone, where most compact, is traversed by small linear cavities, and in several respects resembles the less pure fresh-water limestones of Europe. The concretions not unfrequently become so numerous, that they unite and form a continuous stratum, or even the entire mass.

At Bajada de St^a. Fé, in the Province of Entre Rios, beds of sand, limestone, and clay of different qualities, containing sharks’ teeth and sea-shells, underlie the Pampas deposit. The shells, although numerous, are few in kind. Mr. George B. Sowerby informs me that they appear to belong to one of the less ancient tertiary epochs; they consist of Venus nov. spec. near to V. cancellata; Arca nov. spec. near to A. antiquata; a very large oyster, probably an extinct species; an imperfect specimen of a second species of oyster near to O. edulis; and a Pecten near to P. opercularis. These beds pass upwards into an indurated marl, and this again into the red argillaceous earth of the Pampas, containing the remains of those extinct quadrupeds, which every where characterize that deposit. To the southward of the Plata level plains of an uniform composition, interrupted only at wide intervals by hills of crystalline rock, extend to a distance of about three hundred miles; and to the northward for at least an equal space, and probably much further. As might have been expected from the perfectly level surface, wherever a continuous section is presented on the banks of the great rivers, very slight changes of colour show, that the deposit has been accumulated in strata as horizontal as the land, or as the water-line at the base of the cliffs.

In the province of Banda Oriental (to the N. and N. E. of the Plata), and in part of that of Entre Rios, the land, though very low and level, has a foundation of granitic and other primary rocks. These older formations are partially covered, in most parts, by a reddish earthy mass containing a few small calcareous concretions; while in other parts, they are concealed by more regular strata, of indurated marl passing into limestone, of conglomerates, and ferruginous sandstone. The entire formation probably belongs to the same epoch with that of the Pampas deposit. In the earthy mass, even where it is of little thickness, and where it might readily be mistaken for detritus produced from the underlying granites, remains of large quadrupeds have several times been discovered.

On the shores of the Plata and in the neighbouring districts, proofs of a change of level having taken place between the land and the water within a recent period, may be observed. Both near Monte Video and Colonia del Sacramiento, beds of shells are lying on the beach at the height of several feet above the present tidal action. Near Maldonado I saw estuary shells of recent species embedded in clay, and raised above the level of a neighbouring fresh-water lake.

On the banks of the Parana, a shell identical with, or most closely resembling an estuary species (Potamomya labiata, now living in that part of the Plata, where the water is brackish) is accumulated in great masses, which are found some miles inland, and are elevated several yards above the level of the river. Sir Woodbine Parish, also, has in his possession, shells procured from an extensive formation near Ensenada de Barragan (south of Buenos Ayres), which is quarried for lime. Mr. George Sowerby has examined these fossils, and says the following are identical with living kinds; Voluta colocynthis, Dillwyn: V. angulata, Swainson: Buccinum globulosum, Kiener: a variety of Oliva patula: a Cytheræa closely resembling or identical with C. flexuosa, and a fragment of a second species, probably C. purpurascens; Potamomya labiata; and fragments of oysters. There is, however, a species of Mactra in very great numbers, with which Mr. Sowerby is wholly unacquainted. I may observe that I found recent shells of the first five species inhabiting the coast, a short distance to the southward. Some shelly limestone from the same place, which Sir Woodbine Parish had the kindness to show me, resembles that which I saw at Bajada, and in Banda Oriental. These beds, therefore, probably form parts of the Pampas deposit, and are not merely indicative of the period of its elevation. Nevertheless, on the opposite shores of the Plata, near the mouth of the Uruguay, I found lines of sand dunes, where the Mactra and Cytheræa flexuosa were lying in such quantities on the bare surface, that the inhabitants, by merely sifting the sand, collect them for burning into lime.

After these facts we may feel certain, that at a period not very remote, a great bay occupied the area both of the Pampas and of the lower parts of Banda Oriental. Into this bay the rivers which are now united in the one great stream of the Plata, must formerly have carried down (as happens at the present day) the carcasses of the animals, inhabiting the surrounding countries; and their skeletons would thus become entombed in the estuary mud which was then tranquilly accumulating. Nothing less than a long succession of such accidents can account for the vast number of remains now found buried. As their exposure has invariably been due to the intersection of the plain by the banks of some stream, it is not making an extravagant assertion, to say, that any line whatever drawn across the Pampas would probably cross the skeleton of some extinct animal.

At Bajada, a passage, as I have stated, may be traced upwards from the beds containing marine shells, to the estuary mud with the bones of land animals. In another locality a bed of the same mineralogical nature with the Pampas deposit, underlies clay containing large oysters and other shells, apparently the same with those at Bajada. We may, therefore, conclude that at the period when the Arca, Venus, and Oyster were living, the physical condition of the surrounding country was nearly the same, as at the time when the remains of the mammalia were embedded; and therefore that these shells and the extinct quadrupeds probably either co-existed, or that the interval between their respective existences was, in a geological point of view, extremely short. In this part of South America there is reason to believe that the movements of the land have been so regular, that the period of its elevation may be taken as an element in considering the age of any deposit. The circumstance, therefore, that the beds immediately bordering the Plata, contain very nearly the same species of molluscs, with those now existing in the neighbouring sea, harmonizes perfectly with the more ancient (though really modern) tertiary character of the fossils underlying the Pampas deposit at Bajada, situated at a greater height, and at a considerable distance in the interior. I feel little doubt that the final extinction of the several large quadrupeds of La Plata did not take place, until the time when the sea was peopled with all, or nearly all, its present inhabitants.

Bahia Blanca, situated in latitude 39°, and about 250 miles south of the Plata, constitutes the second district, in which I found the remains of quadrupeds. This large bay is nearly surrounded by very low land, on which successive lines of sand dunes mark in many parts the retreat of the water. At some distance inland a formation of highly indurated marl, passing into limestone, forms an escarpment. Beyond this, rocks of the same character extend over a wide and desolate plain, which rises towards the flanks of the distant mountain of the Sierra de la Ventana, composed of quartz. On the low shores of this bay, only two places occur, where any section of the strata can be seen; and at both of these I found fossil remains.