The Migration of North American Birds (1935)

Part 1

Transcriber Note

Text emphasis denoted by =Bold= and Italics.

+---------------------------------------------------------+ | UNITED STATES | | DEPARTMENT OF AGRICULTURE | | | | CIRCULAR No. 363 | | | +---------------------------------------------------------+ | Washington, D. C. October 1935 | +---------------------------------------------------------+ | | | | | THE MIGRATION | | OF NORTH AMERICAN BIRDS | | | | | | By | | FREDERICK C. LINCOLN | | | | Senior Biologist in Charge, Section of Distribution | | and Migration of Birds Division of Wildlife Research, | | Bureau of Biological Survey | | | | | | [Illustration] | | | | | +---------------------------------------------------------+

For sale by the Superintendent of Documents. Washington, D. C. -- Price 10 cents

CIRCULAR No. 363 OCTOBER 1935 UNITED STATES DEPARTMENT OF AGRICULTURE WASHINGTON, D.C.

THE MIGRATION OF NORTH AMERICAN BIRDS

By Frederick C. Lincoln, senior biologist, in charge Section of Distribution and Migration of Birds, Division of Wildlife Research, Bureau of Biological Survey

CONTENTS

Page

Introduction 1

Mystery of migration 3 Historical accounts 3 Advantages of migration 4 Theories of causes of migration 5

When birds migrate 7 Movements of species and groups 8 Nocturnal and diurnal migration 11

How birds migrate 13 Speed of flight andspeed of migration 13 Altitudes at which birds travel 22 Orientation 23 Segregation during migration 25

Where birds migrate 27 Distances of migration vary 27 Short and undetermined migration 27 Variable migrations wi thin species 28 Fall flights not far south of breeding ranges 29 Long-distance migrations 30

Routes of migration 33 Wide and narrow migration lanes 34 Atlantic oceanic route 39 Atlantic coast route and tributaries 42 Mackenzie Valley-Great Lakes-Mississippi Valley route and tributaries 45 Pacific coast route 47 Pacific oceanic route 49 Arctic routes 52

Evolution of migration routes 52

Vertical migration 55

Vagrant migration 55

Perils of migration 56 Storms 56 Aerial obstructions 57 Exhaustion 59

Influence of the weather on migration 59

Problems of migration 61 Banding studies 61 Movements of residents 62 Migration of the white-throated sparrow 63 Migration of the yellow-billed loon 63

Conclusions 65

Bibliography 66

Index 69

=INTRODUCTION=

Where do the birds go each fall that have nested in our dooryards and frequented the neighboring woods, hills, and marshes? Will the same ones return again to their former haunts next spring? What dangers do they face on their round-trip flight and in their winter homes? These and other questions on the migratory habits of birds puzzle the minds of many who are interested in the feathered species, whether it be the farmer who profits by their tireless warfare against the weed and insect pests of his crops, the bird student who enjoys an abundance and variety of feathered inhabitants about him, or the hunter who wants a continuation from year to year of the sport of wild-fowling. Lack of information on the subject may mean the loss of an important resource by unconsciously letting it slip from us. Ignorance of the facts may be responsible for inadequate legal protection for such species as may urgently need it. More general knowledge on the subject will aid in the perpetuation of the various migrants, the seasonal habitats of some of which are in grave danger from man's utilization, sometimes unwisely, of the marsh, water, and other areas they formerly frequented.

The migrations of birds were probably among the first natural phenomena to attract the attention and intrigue the imagination of man. Recorded observations on the subject date back nearly 3,000 years, to the times of Hesiod, Homer, Herodotus, Aristotle, and others. In the Bible are several references to the periodic movements of birds, as in the book of Job (39:26), where the inquiry is made: "Doth the hawk fly by Thy wisdom and stretch her wings toward the south?" Jeremiah (8:7), wrote: "The stork in the heavens knoweth her appointed time; and the turtle , and the crane, and the swallow, observe the time of their coming." And the flight of quail that saved the Israelites from starvation in their wanderings in the wilderness of Sinai is now recognized as a vast movement of migratory quail (Coturnix coturnix) between their breeding grounds and their winter home in Africa.

Throughout the ages the return flights of migratory birds have been important (1) as a source of food after a lean winter, and (2) as the harbinger of a change in season. The arrival of certain species has been heralded with appropriate ceremonies in many lands, and among Eskimo and other tribes the phenomenon to this day is the accepted sign of the imminence of spring and of warmer weather. The pioneer fur traders in Alaska and Canada offered rewards to the Indian or Eskimo who saw the first goose of the spring, and all joined in jubilant welcome to the newcomer.

Part 2

Always hunted for food, the large flocks of ducks and geese became objects of the enthusiastic attention of an increasing army of sportsmen as the North American Continent became ever more thickly settled. Most of the nongame species were found to be valuable also as allies of the farmer in his never-ending warfare against weed and insect pests. The need for laws protecting the valuable game and nongame birds and for regulated hunting of the diminishing game species followed as a natural course. In the management of this wildlife resource it has become obvious that continuous studies must be made of the food habits of the various species, their environmental needs, and their travels. Hence bird investigations are made by the Biological Survey, the Bureau charged by Congress under the Migratory Bird Treaty Act with the duty of protecting those species that in their yearly journeys pass back and forth between the United States and Canada.

For half a century the Biological Survey has been collecting data on the interesting and important phenomenon of the migration of North American birds. The field men of the Bureau have gathered information concerning the distribution and seasonal movements of the different species in many extended areas, from the Arctic coast south to the pampas of Argentina. Supplementing these investigations is the work of hundreds of volunteer ornithologists and bird students throughout the United States and Canada, who each year, spring and fall, forward to the Bureau reports on migrations observed in their respective localities. Added, to the mass of data thus assembled is a rapidly growing file of records of birds that have been banded and of the subsequent recovery of the marked individuals.

These data, together with other carded records gleaned by the Biological Survey from a vast literature, constitute a series of files that now contain well over 2,500,000 entries, easily the greatest existing accumulation of information pertaining to the distribution and movements of North American birds. Not only do the facts thus assembled form the basis of regulatory action for the protection of the birds, but they also make it possible to publish scientific accounts of the ranges and migrations of the different species. They furnish the basis of this publication.

The several important bird-protective measures adopted by State and Federal Governments, particularly those having as their objective the conservation of the migratory song, insectivorous, and game species, can be effective only if they have intelligent public support. To increase such support, information must be more generally available on that little understood but universally fascinating subject of bird migration. A brief presentation of facts on the migratory habits of the birds, scientifically gathered by the Bureau of Biological Survey over many years, will be helpful to bird-study classes, to conservation organizations, and to farmers and others individually interested in the welfare of the birds.

=THE MYSTERY OF MIGRATION=

HISTORICAL ACCOUNTS

Of observers whose writings are extant, Aristotle, naturalist and philosopher of ancient Greece, was one of the first to discuss the subject of bird migration. He noted that cranes traveled from the steppes of Scythia to the marshes at the headwaters of the Nile, and that pelicans, geese, swans, rails, doves, and many other birds likewise passed to warmer regions to spend the winter. In the earliest years of the Christian era, the elder Pliny, Roman naturalist, in his Historia Naturalis, repeated much of what Aristotle had said on migration, and added comments of his own concerning the movements of the European blackbird, the starling, and the thrushes.

In spite of the keen perception shown in some of his statements, Aristotle also sponsored some superstitions on bird migration that persisted for several centuries. One of these, that of hibernation, became so firmly rooted that in 1878, the American ornithologist Coues (20) listed the titles of no less than 182 papers dealing with the hibernation of swallows (Hirundinidae). The hibernation theory accounted for the autumnal disappearance of certain species of birds as their passing the cold season in a torpid state, hidden in hollow trees, caves, or the mud of marshes. Aristotle ascribed hibernation not only to swallows but also to storks, kites, doves, and others. Some early naturalists wrote fantastic accounts of flocks of swallows seen congregating in the marshes until their accumulated weight bent into the water the reeds on which they clung and thus submerged the birds. It was even recorded that when fishermen in northern waters drew up their nets they sometimes had a mixed "catch" purported to consist of fish and hibernating swallows. Clarke (4) quotes Olaus Magnus, Archbishop of Upsala, who in 1555 published a work entitled "Historia de Gentibus Septentrionalibus et Natura", wherein he observed that if swallows so caught were taken into a warm room they soon begin to fly about but would live only a short time.

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