Part 30

“As the report of the rifle sounded, he plunged madly forward and was instantly lost to sight in the thick scrub. But I felt sure he carried death with him, and so it proved, for we found him lying dead not twenty yards from where he had stood when the bullet struck him. The fatal missile had passed right through his shoulders, and, having expanded on impact, had torn his heart to pieces. I had the dead ewe brought to where the ram had fallen, and laid them side by side; and then stood admiring them for a long time before I could bring myself to skin them. To thus secure a very fine pair of Inyala Antelopes—whose excellently-mounted skins are now safe in the Mammalian Gallery of the new Natural History Museum in Cromwell Road—on the very first day I had hunted for them, and after a little more than an hour’s search, was indeed a most glorious and exceptional piece of good fortune; which, however, has been balanced by many and many a day that I can remember of unrequited labour in search of game.

“As soon as I had stripped the skins, with the leg-bones still attached, from my two beautiful specimens, I had them carried, together with the skulls, to Gugawi’s kraal on the edge of the bush, and there spent the remainder of the day in preparing them for mounting. Of the meat, which was all brought in, I sent a couple of haunches over to Mr. Wissels, and then, after keeping a small piece for myself, gave the remainder to Gugawi to divide amongst his people as he thought fit.

“Next morning I was up and out in the bush just as day was breaking, accompanied only by my guide of yesterday and Longman, who, however, kept some distance behind, in order to allow my guide and myself to approach our game as noiselessly as possible. We had been creeping about in the dense jungle for some three hours without having seen anything, although there was a good deal of fresh spoor about, and twice we had heard Inyalas dash away through the bush without getting a sight of them, when suddenly my guide crouched to the ground, at the same time pointing towards a large ant-heap growing out of the dense scrub, and itself covered with undergrowth. Following the direction of his arm, I made out a reddish patch not fifteen yards away in the gloom of the bush; and taking it for an Inyala ewe, I fired into it point blank, as I required another specimen for mounting. At the shot the animal fell, and, on creeping up to it, I found that it was a young ram. It was something less in size than a full-grown female, from which it did not differ in any way in coloration, and the number and distribution of white stripes and spots. It was thus interesting, as showing that the Inyala changes in general colour from red to grey, only losing the rufous and orange tints on the ears and forehead, which were still conspicuous in the type-specimen described by Mr. Angas, when fully adult.

“On returning to the kraal, Gugawi proposed to take me to a spot some few miles higher up the Usutu, where he said there were plenty of Inyalas, whilst at the same time the bush was not so dense as near his kraal. Being by this time thoroughly sick of crawling about bent nearly double, I hailed with delight the idea of finding the game I was seeking in a country where I could walk upright, and visions of Inyala feeding through open glades passed through my mind; visions, alas! which were never realized, for in my small experience I never found these Antelope anywhere except in dense bush. However, I was glad of the change, and soon had everything ready for a move.

“In the afternoon we travelled some five or six miles up the river, and pitched camp in a bit of jungle near the water’s edge. The Usutu River is here very broad, and reminded me strongly of parts of the Chobi; but whereas the banks of the latter river, as I knew it in the early seventies, abounded in game of many descriptions, from the elephant downwards, there was not a track to be seen along the Usutu of any kind of animal with the exception of the Inyala. All the wealth of wild life which Baldwin saw in this same district forty years ago has melted away before the guns of the native Amatonga hunters; for, be it noted, this is a country in which but very little game has been killed by white men. Rhinoceroses, buffaloes, koodoos, waterbucks, impalas, lions, all are gone—the only game left being the Inyalas, which owe their preservation to the dense jungles in which they live; and even they are being rapidly killed off, as the natives are always after them, lying in wait for them in the paths made by the hippopotami, or creeping stealthily through the bush in their pursuit.

“It would be but tedious reading were I to continue to describe in detail my further bush-crawling experiences in search of Inyalas. Suffice it to say that on Oct. 1 and 2 I secured two more good rams, and preserved their heads for my own Collection. Although I should have liked to get a fourth ram for the South- African Museum, I did not think it prudent to remain any longer in my camp on the edge of a swamp, where I knew the air must be reeking with malarial poison, as, besides the exhalations from the marsh, the ground (from which I was only separated at nights by a little dry grass and a blanket) had been soaked to the depth of 2 feet by the recent rain, thus rendering the conditions more than usually unhealthy. The weather, too, was now again looking very threatening, and I did not relish the idea of any further lying out in the rain; as I knew, from former experience, that I should probably have to pay for the wettings I had already suffered by some attacks of fever—a disease from which I had been entirely exempt for seven years, but the poison of which I knew was still in my blood, and would be likely to be again stirred into activity by my recent exposure to unhealthy conditions.

“Hence, on Saturday, Oct. 3, I packed up my things and returned to Gugawi’s kraal, walking on in the afternoon to Mr. Wissels’s store, and thence to Lourenço Marques, Delagoa Bay, which I reached on October 7th, after a hot and weary tramp.”

Until lately the Inyala was believed to be restricted to the coast-lands of Eastern Africa south of the Zambesi. Recently, however, it has been discovered that this Antelope is likewise found further northward on the Upper Shiré, where it is known to the natives as the “Bō,” the o being pronounced very long. Mr. Alfred Sharpe, C.B., on his return to England at the end of 1891, first brought home a single flat skin of the so-called “Bō,” which was identified by Sclater as belonging to the male of this species, and other specimens have since been obtained in the same district. Mr. Sharpe’s information was that it is found only in a piece of thick scrubby country bordering the Moanza River, which enters the Shiré on its right bank, near the Murchison Cataracts.

In 1895 a fine specimen of this Antelope was forwarded to the British Museum by Mr. Gerald Oliver, R.N., of H.M.S. ‘Herald,’ with the following information in an accompanying letter:—

“On the 5th of October, last year, I was shooting near a village called Mantana’s (lat. 16° 30´ S., long. 35° E.), about 7´ W. by S. of Chilomo, near the right bank of the Shiré River. Impala (Æpyceros melampus) are very plentiful about this particular spot, but I had not been able to get a shot at a good head. Later in the day, wanting meat, I decided to kill what I could, and coming across a solitary doe I fired. Great was the astonishment of myself and boys to find I had killed a female Inyala. I took the skin to Chilomo, and was told it was the first Inyala ever known to have been killed about these parts, and that it was practically an unknown animal there.

“On the 26th of October, near the same spot, a male Inyala was killed by the then Surgeon of this ship. On the 29th of October I saw together 2 Inyalas and 8 Impalas. On the following day, at dusk, I met with and killed a solitary male Inyala, whose skin I forward. The natives were tremendously excited, dancing about and exclaiming ‘Bōōh,’ ‘Bōōh,’ the local name for the Inyala. They examined the body with great care. I was told in the village that the natives, on account of some superstition, would not touch the meat. Our bluejackets, however, ate it, and it was very good. The skin was remarkably glossy and soft, and the hair was long under the neck.”

In 1893 (see P. Z. S. 1893, p. 729) Sclater found a skull of this Antelope in one of Mr. Crawshay’s collections from the district of Lake Mweru, but we are not quite certain that it was actually obtained in that locality. Sir Harry Johnston, in his volume on ‘British Central Africa,’ sums up his information on this species as follows:—

“I am inclined to think that the Inyala Antelope of British Central Africa is limited in its range, so far as we yet know, to the Western and Upper Shiré districts and the Lake Mweru district, and that it may be of a different form from the Inyala of South- east Africa, inasmuch as the males retain the white spots and stripes on the skin to a greater extent, and do not assume such a grey fur at maturity. The Inyala, locally called Bōō, is a very rare animal, frequenting dense thickets. Its horns somewhat resemble those of the Bushbuck, but are much larger proportionately, much wider apart, and slenderer. They may measure as much as 22½ inches in length along the curve (I have a pair of horns giving this measurement). I have only twice seen skins of the adult animal. They were extraordinarily beautiful in colour, the females a deep chestnut, with narrow stripes and spots in pure white, and a black line along the middle of the back from the neck to the base of the tail; the male purplish-grey, with white markings.”

There is also some reason for suspecting that the Inyala, or a nearly allied form, extends even as far north as British East Africa. This suspicion rests upon the evidence supplied by a pair of horns, not specifically distinguishable from those of the typical Tragelaphus angasi, which were procured from the Mau Forest by Mr. F. J. Jackson, C.B. These horns, which are now in the British Museum, were exhibited by Sclater before the Zoological Society of London in May 1897, and are figured on p. 455 of the Society’s ‘Proceedings’ for that year (see fig. 107, p. 147). Unfortunately no skin accompanied the horns; and the specific determination of the Antelope that bore them is rendered uncertain on account of the description of the animal given by the native hunter who killed it. This description, if accurate, certainly applies to a species of Bushbuck distinct both from the Inyala and from all the other known species of Tragelaphus. Hence it is earnestly to be hoped that Mr. Jackson will soon be successful in his attempt to clear up the mystery in which the identity of this interesting Antelope is involved.

We are not aware that any specimen of Angas’ Antelope has ever been brought to Europe alive, and it is by no means commonly met with in our museums. The British Museum contains an adult pair mounted, obtained in St. Lucia Bay by Mr. R. S. Fellowes in 1871, also specimens from the Pongolo River, Zululand, obtained by Mr. Eastwood, and from the Maputa River, obtained by Proudfoot. There are also in the National Collection the specimens from Nyasaland procured by Mr. Sharpe, Sir Harry Johnston, Mr. Crawshay, and Lieut. Oliver, of which we have made mention above. Finally, there is a good mounted pair of specimens obtained by Mr. Selous during his special expedition in quest of this Antelope.

Our principal illustration of Angas’ Antelope (Plate XCII.) was put upon the stone by Mr. Smit, under the directions of the late Sir Victor Brooke, from sketches made by Wolf, and was probably taken from the mounted specimens then in the British Museum. At the same time the woodcut (fig. 106, p. 146), also containing figures of both sexes, was prepared by Mr. Smit.

November, 1899.

Genus III. LIMNOTRAGUS (nom. nov.).

Type.

Hydrotragus, =Gray=, Cat. Rum. B. M. p. 49 (1872) (nec Fitz.) L. spekii.

Medium or large-sized Antelopes allied to Tragelaphus, but with rougher and shaggier coats, longer legs, and with horns which more nearly approach those of Strepsiceros, as they show a strong tendency to assume a third twist. Further, Limnotragus differs strikingly from Tragelaphus and Strepsiceros in the structure of its feet, the hoofs being often nearly three times as long (measured along the front edge) as thick (measured along the margin of the pastern). Moreover, the skin which covers the back of the pastern is denuded of hair, and thick and horny, being practically of the same consistency as the upper rim of the posterior side of the hoof.

It is not without some hesitation that we refer to a separate genus those species of Tragelaphine Antelopes (hitherto placed in Tragelaphus) which have undergone certain special modifications of structure in adaptation to a semi-aquatic mode of life. In the species of Tragelaphus discussed in the preceding part of this work, as well as in all the other genera of Tragelaphinæ, the feet adhere to the digitigrade type characteristic of most of the ruminant artiodactyle Ungulates, retaining the short narrow hoofs and strong elastic ankles fitted for easy and swift progress over the firm soil of the veldt or woodland. In the species of Limnotragus, on the contrary, the feet are furnished with hoofs of relatively enormous length, which spread far apart at every step, and are obviously designed to enable their owner to pass over the soft soil of marshes and river-banks without sinking deeply into the ground. This modification is accompanied by an increase in the flexibility of the ankle-joints, which are capable of yielding to the weight of the body, so as to allow the false-hoofs and the smooth tough horny skin at the back of the pasterns to rest upon the soil, and thus to further augment the supporting area of the foot. It is these peculiarities in the structure of the feet, in conjunction with a length of limb exceeding that of other Tragelaphines, which impart to the species of Limnotragus that characteristic semiplantigrade aspect and that unusual awkwardness of gait so noticeable in these Antelopes when walking upon firm resisting ground.

Up to the present time three species referable to this genus have been described, namely, L. spekii from E. Africa, L. gratus from Tropical West Africa, and L. selousi from the valley of the Zambesi. Unfortunately there are not at present available materials from the different parts of the area over which the genus ranges, sufficient to enable us to determine satisfactorily the exact value to be assigned to these three forms. Consequently, although the evidence, so far as it goes, tends to show that the characters upon which they have been based may ultimately prove to have merely a subspecific importance, we prefer, for the time being, to allow them to take the rank that was originally assigned to them by their respective describers, and to arrange them as three species.

Range of the Genus. Congo Valley and Lake-districts of Southern and Eastern Africa.

The three species may be shortly distinguished as follows:—

a. Size smaller; height at withers about 36 inches: sexes dissimilar; male blackish, female rufous 127. L. spekii. b. Size larger; height at withers about 40 inches. a1. Sexes similar, blackish 128. L. selousi. b1. Sexes dissimilar; male blackish brown, female red. 129. L. gratus.

127. SPEKE’S SITATUNGA.

LIMNOTRAGUS SPEKII (Sclater).