Part 4

Although little acquainted with biological science, Whewell seems to have taken particular pains with that part of his work which deals with the history of geological and biological speculation; and several chapters of his seventeenth and eighteenth books, which comprise the history of physiology, of comparative anatomy and of the palætiological sciences, vividly reproduce the controversies of the early days of the Victorian epoch. But here, as in the case of the doctrine of the conservation of energy, the historian of the inductive sciences has no prophetic insight; not even a suspicion of that which the near future was to bring forth. And those who still repeat the once favorite objection that Darwin's 'Origin of Species' is nothing but a new version of the 'Philosophie zoologique' will find that, so late as 1844, Whewell had not the slightest suspicion of Darwin's main theorem, even as a logical possibility. In fact, the publication of that theorem by Darwin and Wallace, in 1859, took all the biological world by surprise. Neither those who were inclined towards the 'progressive transmutation' or 'development' doctrine, as it was then called, nor those who were opposed to it, had the slightest suspicion that the tendency to variation in living beings, which all admitted as a matter of fact; the selective influence of conditions, which no one could deny to be a matter of fact, when his attention was drawn to the evidence; and the occurrence of great geological changes which also was matter of fact; could be used as the only necessary postulates of a theory of the evolution of plants and animals which, even if not at once, competent to explain all the known facts of biological science, could not be shown to be inconsistent with any. So far as biology is concerned, the publication of the 'Origin of Species,' for the first time, put the doctrine of evolution, in its application to living things, upon a sound scientific foundation. It became an instrument of investigation, and in no hands did it prove more brilliantly profitable than in those of Darwin himself. His publications on the effects of domestication in plants and animals, on the influence of cross-fertilisation, on flowers as organs for effecting such fertilisation, on insectivorous plants, on the motions of plants, pointed out the routes of exploration which have since been followed by hosts of inquirers, to the great profit of science.

Darwin found the biological world a more than sufficient field for even his great powers, and left the cosmical part of the doctrine to others. Not much has been added to the nebular hypothesis, since the time of Laplace, except that the attempt to show (against that hypothesis) that all nebulæ are star clusters, has been met by the spectroscopic proof of the gaseous condition of some of them. Moreover, physicists of the present generation appear now to accept the secular cooling of the earth, which is one of the corollaries of that hypothesis. In fact, attempts have been made, by the help of deductions from the data of physics, to lay down an approximate limit to the number of millions of years which have elapsed since the earth was habitable by living beings. If the conclusions thus reached should stand the test of further investigation, they will undoubtedly be very valuable. But, whether true or false, they can have no influence upon the doctrine of evolution in its application to living organisms. The occurrence of successive forms of life upon our globe is an historical fact, which cannot be disputed; and the relation of these successive forms, as stages of evolution of the same type, is established in various cases. The biologist has no means of determining the time over which the process of evolution has extended, but accepts the computation of the physical geologist and the physicist, whatever that may be.

Evolution as a philosophical doctrine applicable to all phenomena, whether physical or mental, whether manifested by material atoms or by men in society, has been dealt with systematically in the 'Synthetic Philosophy' of Mr. Herbert Spencer. Comment on that great undertaking would not be in place here. I mention it because, so far as I know, it is the first attempt to deal, on scientific principles, with modern scientific facts and speculations. For the 'Philosophic positive' of M. Comte, with which Mr. Spencer's system of philosophy is sometimes compared, though it professes a similar object, is unfortunately permeated by a thoroughly unscientific spirit, and its author had no adequate acquaintance with the physical sciences even of his own time.

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The doctrine of evolution, so far as the present physical cosmos is concerned, postulates the fixity of the rules of operation of the causes of motion in the material universe. If all kinds of matter are modifications of one kind, and if all modes of motion are derived from the same energy, the orderly evolution of physical nature out of one substratum and one energy implies that the rules of action of that energy should be fixed and definite. In the past history of the universe, back to that point, there can be no room for chance or disorder. But it is possible to raise the question whether this universe of simplest matter and definitely operating energy, which forms our hypothetical starting point, may not itself be a product of evolution from a universe of such matter, in which the manifestations of energy were not definite--in which, for example, our laws of motion held good for some units and not for others, or for the same units at one time and not at another--and which would therefore be a real epicurean chance-world?

For myself, I must confess that I find the air of this region of speculation too rarefied for my constitution, and I am disposed to take refuge in 'ignoramus et ignorabimus.'

The execution of my further task, the indication of the most important achievements in the several branches of physical science during the last fifty years, is embarrassed by the abundance of the objects of choice; and by the difficulty which everyone, but a specialist in each department, must find in drawing a due distinction between discoveries which strike the imagination by their novelty, or by their practical influence, and those unobtrusive but pregnant observations and experiments in which the germs of the great things of the future really lie. Moreover, my limits restrict me to little more than a bare chronicle of the events which I have to notice.

In physics and chemistry, the old boundaries of which sciences are rapidly becoming effaced, one can hardly go wrong in ascribing a primary value to the investigations into the relation between the solid, liquid, and gaseous states of matter on the one hand, and degrees of pressure and of heat on the other. Almost all, even the most refractory, solids have been vaporised by the intense heat of the electric arc; and the most refractory gases have been forced to assume the liquid, and even the solid, forms by the combination of high pressure with intense cold. It has further been shown that there is no discontinuity between these states--that a gas passes into the liquid state through a condition which is neither one nor the other, and that a liquid body becomes solid, or a solid liquid, by the intermediation of a condition in which it is neither truly solid nor truly liquid.

Theoretical and experimental investigations have concurred in the establishment of the view that a gas is a body, the particles of which are in incessant rectilinear motion at high velocities, colliding with one another and bounding back when they strike the walls of the containing vessel; and, on this theory, the already ascertained relations of gaseous bodies to heat and pressure have been shown to be deducible from mechanical principles. Immense improvements have been effected, in the means of exhausting a given space of its gaseous contents; and experimentation on the phenomena which attend the electric discharge and the action of radiant heat, within the extremely rarefied media thus produced, has yielded a great number of remarkable results, some of which have been made familiar to the public by the Gieseler tubes and the radiometer. Already, these investigations have afforded an unexpected insight into the constitution of matter and its relations with thermal and electric energy, and they open up a vast field for future inquiry into some of the deepest problems of physics. Other important steps, in the same direction, have been effected by investigations into the absorption of radiant heat proceeding from different sources by solid, fluid, and gaseous bodies. And it is a curious example of the interconnection of the various branches of physical science, that some of the results thus obtained have proved of great importance in meteorology.

The existence of numerous dark lines, constant in their number and position in the various regions of the solar spectrum, was made out by Fraunhofer in the early part of the present century, but more than forty years elapsed before their causes were ascertained and their importance recognised. Spectroscopy, which then took its rise, is probably that employment of physical knowledge, already won, as a means of further acquisition, which most impresses the imagination. For it has suddenly and immensely enlarged our power of overcoming the obstacles which almost infinite minuteness on the one hand, and almost infinite distance on the other, have hitherto opposed to the recognition of the presence and the condition of matter. One eighteen-millionth of a grain of sodium in the flame of a spirit-lamp may be detected by this instrument; and, at the same time, it gives trust-worthy indications of the material constitution not only of the sun, but of the farthest of those fixed stars and nebulæ which afford sufficient light to affect the eye, or the photographic plate, of the inquirer.

The mathematical and experimental elucidation of the phenomena of electricity, and the study of the relations of this form of energy with chemical and thermal action, had made extensive progress before 1837. But the determination of the influence of magnetism on light, the discovery of diamagnetism, of the influence of crystalline structure on magnetism, and the completion of the mathematical theory of electricity, all belong to the present epoch. To it also appertain the practical execution and the working out of the results of the great international system of observations on terrestrial magnetism, suggested by Humboldt in 1836; and the invention of instruments of infinite delicacy and precision for the quantitative determination of electrical phenomena. The voltaic battery has received vast improvements; while the invention of magneto-electric engines and of improved means of producing ordinary electricity has provided sources of electrical energy vastly superior to any before extant in power, and far more convenient for use.

It is perhaps this branch of physical science which may claim the palm for its practical fruits, no less than for the aid which it has furnished to the investigation of other parts of the field of physical science. The idea of the practicability of establishing a communication between distant points, by means of electricity, could hardly fail to have simmered in the minds of ingenious men since, well nigh a century ago, experimental proof was given that electric disturbances could be propagated through a wire twelve thousand feet long. Various methods of carrying the suggestion into practice had been carried out with some degree of success; but the system of electric telegraphy, which, at the present time, brings all parts of the civilised world within a few minutes of one another, originated only about the commencement of the epoch under consideration. In its influence on the course of human affairs, this invention takes its place beside that of gunpowder, which tended to abolish the physical inequalities of fighting men; of printing, which tended to destroy the effect of inequalities in wealth among learning men; of steam transport, which has done the like for travelling men. All these gifts of science are aids in the process of levelling up; of removing the ignorant and baneful prejudices of nation against nation, province against province, and class against class; of assuring that social order which is the foundation of progress, which has redeemed Europe from barbarism, and against which one is glad to think that those who, in our time, are employing themselves in fanning the embers of ancient wrong, in setting class against class, and in trying to tear asunder the existing bonds of unity, are undertaking a futile struggle. The telephone is only second in practical importance to the electric telegraph. Invented, as it were, only the other day, it has already taken its place as an appliance of daily life. Sixty years ago, the extraction of metals from their solutions, by the electric current, was simply a highly interesting scientific fact. At the present day, the galvano-plastic art is a great industry; and, in combination with photography, promises to be of endless service in the arts. Electric lighting is another great gift of science to civilisation, the practical effects of which have not yet been fully developed, largely on account of its cost. But those whose memories go back to the tinder-box period, and recollect the cost of the first lucifer matches, will not despair of the results of the application of science and ingenuity to the cheap production of anything for which there is a large demand.

The influence of the progress of electrical knowledge and invention upon that of investigation in other fields of science is highly remarkable. The combination of electrical with mechanical contrivances has produced instruments by which, not only may extremely small intervals of time be exactly measured, but the varying rapidity of movements, which take place in such intervals and appear to the ordinary sense instantaneous, is recorded. The duration of the winking of an eye is a proverbial expression for an instantaneous action; but, by the help of the revolving cylinder and the electrical marking-apparatus, it is possible to obtain a graphic record of such an action, in which, if it endures a second, that second shall be subdivided into a hundred, or a thousand, equal parts, and the state of the action at each hundredth, or thousandth, of a second exhibited. In fact, these instruments may be said to be time-microscopes. Such appliances have not only effected a revolution in physiology, by the power of analysing the phenomena of muscular and nervous activity which they have conferred, but they have furnished new methods of measuring the rate of movement of projectiles to the artillerist. Again, the microphone, which renders the minutest movements audible, and which enables a listener to hear the footfall of a fly, has equipped the sense of hearing with the means of entering almost as deeply into the penetralia of nature, as does the sense of sight.

That light exerts a remarkable influence in bringing about certain chemical combinations and decompositions was well known fifty years ago, and various more or less successful attempts to produce permanent pictures, by the help of that knowledge, had already been made. It was not till 1839, however, that practical success was obtained; but the 'daguerreotypes' were both cumbrous and costly, and photography would never have attained its present important development had not the progress of invention substituted paper and glass for the silvered plates then in use. It is not my affair to dwell upon the practical application of the photography of the present day, but it is germane to my purpose to remark that it has furnished a most valuable accessory to the methods of recording motions and lapse of time already in existence. In the hands of the astronomer and the meteorologist, it has yielded means of registering terrestrial, solar, planetary, and stellar phenomena, independent of the sources of error attendant on ordinary observation; in the hands of the physicist, not only does it record spectroscopic phenomena with unsurpassable ease and precision, but it has revealed the existence of rays having powerful chemical energy, or beyond the visible limits of either end of the spectrum; while, to the naturalist, it furnishes the means by which the forms of many highly complicated objects may be represented, without that possibility of error which is inherent in the work of the draughtsman. In fact, in many cases, the stern impartiality of photography is an objection to its employment: it makes no distinction between the important and the unimportant; and hence photographs of dissections, for example, are rarely so useful as the work of a draughtsman who is at once accurate and intelligent.

The determination of the existence of a new planet, Neptune, far beyond the previously known bounds of the solar system, by mathematical deduction from the facts of perturbation; and the immediate confirmation of that determination, in the year 1846, by observers who turned their telescopes into the part of the heavens indicated as its place, constitute a remarkable testimony of nature to the validity of the principles of the astronomy of our time. In addition, so many new asteroids have been added to those which were already known to circulate in the place which theoretically should be occupied by a planet, between Mars and Jupiter, that their number now amounts to between two and three hundred. I have already alluded to the extension of our knowledge of the nature of the heavenly bodies by the employment of spectroscopy. It has not only thrown wonderful light upon the physical and chemical constitution of the sun, fixed stars, and nebulæ, and comets, but it holds out a prospect of obtaining definite evidence as to the nature of our so-called elementary bodies.

The application of the generalisations of thermotics to the problem of the duration of the earth, and of deductions from tidal phenomena to the determination of the length of the day and of the time of revolution of the moon, in past epochs of the history of the universe; and the demonstration of the competency of the great secular changes, known under the general name of the precession of the equinoxes, to cause corresponding modifications in the climate of the two hemispheres of our globe, have brought astronomy into intimate relation with geology. Geology, in fact, proves that, in the course of the past history of the earth, the climatic conditions of the same region have been widely different, and seeks the explanation of this important truth from the sister sciences. The facts that, in the middle of the Tertiary epoch, evergreen trees abounded within the arctic circle; and that, in the long subsequent Quaternary epoch, an arctic climate, with its accompaniment of gigantic glaciers, obtained in the northern hemisphere, as far south as Switzerland and Central France, are as well established as any truths of science. But, whether the explanation of these extreme variations in the mean temperature of a great part of the northern hemisphere is to be sought in the concomitant changes in the distribution of land and water surfaces of which geology affords evidence, or in astronomical conditions, such as those to which I have referred, is a question which must await its answer from the science of the future.

Turning now to the great steps in that progress which the biological sciences have made since 1837, we are met, on the threshold of our epoch, with perhaps the greatest of all--namely, the promulgation by Schwann, in 1839, of the generalisation known as the 'cell theory,' the application and extension of which by a host of subsequent investigators has revolutionised morphology, development, and physiology. Thanks to the immense series of labors thus inaugurated, the following fundamental truths have been established.

All living bodies contain substances of closely similar physical and chemical composition, which constitute the physical basis of life, known as protoplasm. So far as our present knowledge goes, this takes its origin only from pre-existing protoplasm.

All complex living bodies consist, at one period of their existence, of an aggregate of minute portions of such substance, of similar structure, called cells, each cell having its own life independent of the others, though influenced by them.

All the morphological characters of animals and plants are the results of the mode of multiplication, growth, and structural metamorphosis of these cells, considered as morphological units.

All the physiological activities of animals and plants--assimilation, secretion, excretion, motion, generation--are the expression of the activities of the cells considered as physiological units. Each individual, among the higher animals and plants, is a synthesis of millions of subordinate individualities. Its individuality, therefore, is that of a 'civitas' in the ancient sense, or that of the Leviathan of Hobbes.

There is no absolute line of demarcation between animals and plants. The intimate structure, and the modes of change, in the cells of the two are fundamentally the same. Moreover, the higher forms are evolved from lower, in the course of their development, by analogous processes of differentiation, coalescence, and reduction in both the vegetable and the animal worlds.

At the present time, the cell theory, in consequence of recent investigations into the structure and metamorphosis of the 'nucleus,' is undergoing a new development of great significance, which, among other things, foreshadows the possibility of the establishment of a physical theory of heredity, on a safer foundation than those which Buffon and Darwin have devised.

The popular belief in abiogenesis, or the so-called 'spontaneous' generation of the lower forms of life, which was accepted by all the philosophers of antiquity, held its ground down to the middle of the seventeenth century. Notwithstanding the frequent citation of the phrase, wrongfully attributed to Harvey, 'Omne vivum ex ovo,' that great physiologist believed in spontaneous generation as firmly as Aristotle did. And it was only in the latter part of the seventeenth century, that Redi, by simple and well-devised experiments, demonstrated that, in a great number of cases of supposed spontaneous generation, the animals which made their appearance owed their origin to the ordinary process of reproduction, and thus shook the ancient doctrine to its foundations. In the middle of the eighteenth century, it was revived, in a new form, by Needham and Buffon; but the experiments of Spallanzani enforced the conclusions of Redi, and compelled the advocates of the occurrence of spontaneous generation to seek evidence for their hypothesis only among the parasites and the lowest and minutest organisms. It is just fifty years since Schwann and others proved that, even with respect to them, the supposed evidence of abiogenesis was untrustworthy.

During the present epoch, the question, whether living matter can be produced in any other way than by the physiological activity of other living matter, has been discussed afresh with great vigor; and the problem has been investigated by experimental methods of a precision and refinement unknown to previous investigators. The result is that the evidence in favor of abiogenesis has utterly broken down, in every case which has been properly tested. So far as the lowest and minutest organisms are concerned, it has been proved that they never make their appearance, if those precautions by which their germs are certainly excluded are taken. And, in regard to parasites, every case which seemed to make for their generation from the substance of the animal, or plant, which they infest has been proved to have a totally different significance. Whether not-living matter may pass, or ever has, under any conditions, passed into living matter, without the agency of pre-existing living matter, necessarily remains an open question; all that can be said is that it does not undergo this metamorphosis under any known conditions. Those who take a monistic view of the physical world may fairly hold abiogenesis as a pious opinion, supported by analogy and defended by our ignorance. But, as matters stand, it is equally justifiable to regard the physical world as a sort of dual monarchy. The kingdoms of living matter and of not-living matter are under one system of laws, and there is a perfect freedom of exchange and transit from one to the other. But no claim to biological nationality is valid except birth.

In the department of anatomy and development, a host of accurate and patient inquirers, aided by novel methods of preparation, which enable the anatomist to exhaust the details of visible structure and to reproduce them with geometrical precision, have investigated every important group of living animals and plants, no less than the fossil relics of former faunæ and floræ. An enormous addition has thus been made to our knowledge, especially of the lower forms of life, and it may be said that morphology, however inexhaustible in detail, is complete in its broad features. Classification, which is merely a convenient summary expression of morphological facts, has undergone a corresponding improvement. The breaks which formerly separated our groups from one another, as animals from plants, vertebrates from invertebrates, cryptogams from phanerogams, have either been filled up, or shown to have no theoretical significance. The question of the position of man, as an animal, has given rise to much disputation, with the result of proving that there is no anatomical or developmental character by which he is more widely distinguished from the group of animals most nearly allied to him, than they are from one another. In fact, in this particular, the classification of Linnæus has been proved to be more in accordance with the facts than those of most of his successors.

The study of man, as a genus and species of the animal world, conducted with reference to no other considerations than those which would be admitted by the investigator of any other form of animal life, has given rise to a special branch of biology, known, as Anthropology, which has grown with great rapidity. Numerous societies devoted to this portion of science have sprung up, and the energy of its devotees has produced a copious literature. The physical characters of the various races of men have been studied with a minuteness and accuracy heretofore unknown; and demonstrative evidence of the existence of human contemporaries of the extinct animals of the latest geological epoch has been obtained, physical science has thus been brought into the closest relation with history and with archæology; and the striking investigations which, during our time, have put beyond doubt the vast antiquity of Babylonian and Egyptian civilisation, are in perfect harmony with the conclusions of anthropology as to the antiquity of the human species.

Classification is a logical process which consists in putting together those things which are like and keeping asunder those which are unlike; and a morphological classification, of course, takes notes only of morphological likeness and unlikeness. So long, therefore, as our morphological knowledge was almost wholly confined to anatomy, the characters of groups were solely anatomical; but as the phenomena of embryology were explored, the likeness and unlikeness of individual development had to be taken into account; and, at present, the study of ancestral evolution introduces a new element of likeness and unlikeness which is not only eminently deserving of recognition, but must ultimately predominate over all others. A classification which shall represent the process of ancestral evolution is, in fact, the end which the labors of the philosophical taxonomist must keep in view. But it is an end which cannot be attained until the progress of palæontology has given us far more insight than we yet possess, into the historical facts of the case. Much of the speculative 'phylogeny,' which abounds among my present contemporaries, reminds me very forcibly of the speculative morphology, unchecked by a knowledge of development, which was rife in my youth. As hypothesis, suggesting inquiry in this or that direction, it is often extremely useful; but, when the product of such speculation is placed on a level with those generalisations of morphological truths which are represented by the definitions of natural groups, it tends to confuse fancy with fact and to create mere confusion. We are in danger of drifting into a new 'Natur-Philosophie' worse than the old, because there is less excuse for it. Boyle did great service to science by his 'Sceptical Chemist,' and I am inclined to think that, at the present day, a 'Sceptical Biologist' might exert an equally beneficent influence.