Part 30

G. Massee, the British mycologist, makes of the black-spored and the purple and purplish-brown spored series two divisions, calling them, respectively, Porphyrosporeae and Melanosporeae.

The recorded edible species of the spore section Dermini are found in Pholiota, Cortinarius, and Paxillus. The larger proportion of the Pholiotas grow upon tree stumps. They have a fugacious, persistent friable ring, and are liable to be confused with the Cortinarii, unless attention is paid to the spidery veil and the iron-rust tint of the spores of the latter. Only a few of the species are recorded as edible, but none are known to be poisonous. Cortinarius is a large genus. It contains a larger proportion of edible species than Pholiota, and none are recorded as poisonous. The cobweb-like veil which extends from stem to margin of cap in the young species, and the rust-colored spores which dust the gills as the species mature, distinguish the genus from all others.

A characteristic feature of Paxillus, and one which makes it easily distinguishable from others of the same group, is the ease with which the gills as a whole can be separated from the substance or fleshy portion of the cap. There is an exception to this in the species Paxillus involutus, recorded by Peck as edible.

POLYPOREI.

Hymenium lining the cavity of tubes or pores which are sometimes broken up into teeth or concentric plates.--Berkeley's Outlines.

The plants of this second primary group or order of the family Hymenomycetes exhibit a greater dissimilarity of form and texture than do those of the Agaricini. Some of its genera consist almost wholly of coriaceous or woody plants. A few contain fleshy ones. Some of the species have a distinct stem, while others are stemless. With regard to the receptacle in the plants of the genera Boletus, Strobilomyces, etc., it forms a perfect cap, like that of the common Agaric, a cushion of tubes taking the place of gills on the under surface of the cap, the hymenium in this case lining the inner surface of the tubes from which the spores drop when mature.

In some species, such as those of the genus Poria, the receptacle is reduced to a single thin fibrous stratum, adhering closely to the matrix and exposing a surface of crowded pores, and in others it consists of fibrous strata formed in concentric layers.

A number of groups, each of which was treated in the original Friesian classification as a single genus, have more recently been recognized as comprising several distinct genera. In the Saccardian system the genera Trametes, Daedalea, Merulius, Porothelium, and Fistulina still retain the generic rank assigned to them by Fries, but the old genus Boletus is subdivided into four genera, Boletus, Strobilomyces, Boletinus, and Gyrodon, while Polyporus, originally a very large genus, is subdivided into the genera Polyporus, Fomes, Polystictus, and Poria. This arrangement was in part suggested by Fries in his later works, and is accepted by M. C. Cooke, as indicated in his latest work on fungi.

Quoting M. C. Cooke, "Strobilomyces is Boletus with a rough warty and scaly pileus; Boletinus is Boletus with short, large radiating pores; and Gyrodon is Boletus with elongated sinuate irregular pores, all fleshy, firm fungi of robust habit, possessing stem and cap." The species of the genus Polyporus as now restricted are somewhat fleshy in the young stage, shrinking as they mature and dry, and becoming indurated with age. In Fomes the species, of woody consistency from the first, have no room for shrinkage, and are quite rigid; the tubes being in strata, and the strata growing yearly, the species are virtually perennial. The pileus of the plant shows a rigid polished crust resulting from resinous exudations.

In Polystictus the plants are usually small, thin, tough, and irregular in outline, the tubes exceedingly short, with thin walls, which easily split up, giving the pores at times a toothed or fringed appearance. The surface is velvety, or hairy, and zoned in varying colors. They are very common upon decaying tree stumps, often covering the surface of the stump in gaily colored layers. Not esculent.

Poria is composed of resupinate species with the pores normally in a single series, the whole stratum spread over, and adhering to the matrix. The species are coriaceous or woody. Not esculent.

The plants of the genus Trametes allied to Fomes are epiphytal, with the trama the same in substance and color as the hymenophore. The tubes do not form in regular strata, but are sunk into the substance of the pileus. The plants are coriaceous, and none are edible.

Daedalea closely resembles Trametes with the tubes forming deep labyrinthiform depressions. Whole plant woody, sessile.

Hexagonia, allied by its characteristics to Polystictus, has large hexagonal pores, with firm, entire dissepiments.

In Favolus the plants are slightly fleshy and substipitate with the pores angular, and radiating from the stem. Not edible.

The species of the genus Laschia are recognized by the shallow irregular pores and the vein like character of their dissepiments (or pore walls). Substance slightly gelatinous.

In the plants of Porothelium, irregular papillae take the place of tubes, and the plants are sub-membranaceous and resupinate, having the habit of those of Poria.

The genus Merulius has been termed the lowest and most imperfect of the genera of Polyporei. It presents a soft, waxy spore-bearing surface, reticulated with obtuse folds. Solenia, by early authors placed in Discomycetes, thence transferred to Auricularini, and by some authors associated with Cyphella in Theleporei, now finds place as one of the genera of Polyporei as given by Saccardo.

The above-mentioned genera, together with Myriadoporus, Ceriomyces, Bresadolia, Theleporus, Gloeporus, and Cyclomyces, constitute the Polyporeae of the Saccardian system.

Myriadoporus is a North American genus. It is a form of the genus Polyporus, but with pores in the interior as well as on the exterior surface. Ceriomyces is generally regarded as a spurious genus. It is similar to Myriadoporus, but with internal pores and only spurious pores externally. Of Bresadolia Cooke says "there is only one described species, and of this only one specimen has been found." Theleporus is an African genus of which only one species is known. Gloeporus is a form of resupinate Polyporus, except that the hymenium or pore-bearing surface is gelatinous instead of being firm. Cyclomyces is a genus with some features of Lenzites; it is leathery. All of these are more or less coriaceous. None are edible. Campbellia is a new genus. It is Merulius with a pileus and central stem.

The edible Polyporeae are found in the genera Boletus, Strobilomyces, Gyrodon, Boletinus, Polyporus, and Fistulina. Of these, the first four genera contain most of the edible species as well as a few which have been regarded as unwholesome or poisonous.

In the genus Polyporus as now restricted, the species Polyporus sulphureus Fries is perhaps the one most likely to be selected for table use, the others becoming very quickly indurated or tough, and this should be gathered when very young, as in maturity it loses its fleshy consistency and becomes dry and tough. It is common on old tree stumps and is often found on the dead wood of living trees, the bright yellow and vivid orange red tints which characterize the young plant making it very conspicuous.

It is easily recognized by its irregular, closely overlapping frond-like caps, white flesh, and the very small sulphur-yellow tubes. The spores are white, elliptical. The flesh of young specimens is somewhat juicy.

The geographical distribution is wide, and in places where a moist, warm temperature prevails plants of this species often attain very large proportions, sometimes completely encircling the trunk of a tree at its base. The bright colors fade as the plant matures, and the plant becomes indurated and friable, when very old crumbling readily in the hands.

To prepare for the table, very thin slices of young specimens should be cut and either allowed to slowly simmer on the back of the range, or soaked in milk and then fried in butter.

Of the genus Fistulina but one species, Fistulina hepatica, figured in Plate X, is recorded as edible and indigenous to this country.

PLATE X.

=Fistulina hepatica= Bull. "Beefsteak Mushroom," "Liver Fungus."