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Speciation of the Wandering Shrew · James S. Findley — chapter 8 of 21 · ~874 words · public domain

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Remarks.--S. v. setosus lives mostly in forests. According to Dalquest (1948:139) it is commonest at high altitudes in western Washington. In the Hudsonian Life-zone where shrew habitat is more restricted and marginal than it is at lower altitudes in the humid part of Washington, setosus might be expected to compete with S. v. vagrans and to supplant it. Records of occurrence in the Olympic Mountains suggest a degree of such separation there.

Specimens examined.--Total number, 135.

BRITISH COLUMBIA: Lund, Malaspina Inlet, 4 BS; Gibson's Landing, 10 BS; Port Moody, 19 BS; Langley, 2 BS; Chilliwack, 1 BS; Manning Park, 2 PMBC.

WASHINGTON: Whatcom Co.: Mt. Baker, 6 WSC; Barron, 1 BS. Chelan Co.: Clovay Pass, 1 WSC; Stehekin, 6 (4 WSC, 2 BS); Cascade Tunnel, 1 WSC. King Co.: Scenic, 1 WSC. Kittitas Co.: Lake Kachess, 1 WSC; Easton, 10 BS. Clallam Co.: 8 mi. W Sekin River, 1 WSC; mouth of Sekin River, 1 WSC; Clallam Bay, 1 CMNH; 7 mi. W Port Angeles, 1 WSC; Ozette Lake, 1 CMNH; 12 mi. S Port Angeles, 4 WSC; Forks, 1 CMNH; Deer Lake, 7 CMNH; Hoh Lake, 1 CMNH; Bogachiel Peak, 1 CMNH; Sol Duc Hot Springs, 3 CMNH; Sol Duc Park, 1 CMNH; Canyon Creek, 1 WSC; Sol Duc Divide, 2 WSC; Cat Creek, 2 WSC. Jefferson Co.: Jackson Ranger Station, 1 CMNH; Mt. Kimta, 2 CMNH; Reflection Lake, 6 CMNH; Blue Glacier, 3 CMNH. Gray's Harbor Co.: Westport, 1 WSC. Pierce Co.: Fort Lewis, 1 FC; Mt. Rainier, 19 (16 BS, 3 WSC). Pacific Co.: Tokeland, 2 BS. Yakima Co.: Gotchen Creek, 3 WSC; Mt. Adams, 1 WSC. Skamania Co.: Mt. St. Helens, 1.

OREGON: Hood River Co.: 2 mi. W Parkdale, 2 BS.

Marginal Records.--BRITISH COLUMBIA: Rivers Inlet (Anderson, 1947:20); Agassiz (Jackson, 1928:136); Chilliwack Lake. WASHINGTON: Barron; Lyman Lake (Jackson, 1928:137); Mt. Stuart (Dalquest, 1948:141); Mt. Adams. OREGON: 2 mi. W Parkdale. WASHINGTON: Ilwaco (Jackson, 1928:137); Lund, Malaspina Inlet.

=Sorex vagrans longicauda= Merriam

Sorex obscurus longicauda Merriam, N. Amer. Fauna, 10:74, December 31, 1895.

Type.--Adult male, skin and skull; No. 74711, U. S. Biol. Surv. Coll.; obtained on September 9, 1895 by C. P. Streator, from Wrangell, Alaska.

Range.--The British Columbian and Alaskan coasts from Rivers Inlet north to near Juneau and also certain islands including Etolin, Gravina, Revillagigedo, Sergeif, and Wrangell.

Diagnosis.--Size medium for the species, tail relatively long; average and extreme measurements of 17 specimens from the type locality are: total length, 128.4 (122-138); tail, 57.8 (53-66); hind foot, 15.1 (14-16). Color dark in summer and winter.

Comparisons.--For comparison with S. v. setosus see account of that subspecies. Larger and darker than S. v. obscurus with which it intergrades east of the humid coastal region; larger and darker than S. v. alascensis with which it intergrades in the Lynn Canal area; larger and darker than S. v. calvertensis which occurs on Calvert Island and Banks Island, British Columbia; differs from S. v. insularis of Smythe, Townsend, and Reginald islands in larger size and blackish rather than brown winter pelage; larger and relatively longer-tailed than S. v. elassodon which occurs on most of the islands west of the range of longicauda; larger and relatively longer-tailed than S. v. isolatus.

Specimens examined.--Total number, 151.

ALASKA: Wrangell, 54 BS; 8 AMNH; Crittenden Creek, 1 BS; Ketchikan, 2 BS; Loring, 11 BS.

BRITISH COLUMBIA: Port Simpson, 25 BS; Inverness, 15 BS; head of Rivers Inlet, 35 BS.

Marginal Records.--BRITISH COLUMBIA: Great Glacier, Stikine River (Jackson, 1928:133). ALASKA: Burroughs Bay (ibid.). BRITISH COLUMBIA: Bella Coola region (Anderson, 1947:19); head of Rivers Inlet; Spider Island (Cowan, 1941:101); Goose Island (Cowan, 1941:99); Princess Royal Island (Cowan, 1941:98); Pitt Island (ibid.); Metlakatla (Jackson, 1928:133); Port Simpson. ALASKA: Gravina Island (ibid.); Helm Bay (ibid.); Etolin Island (ibid.); Sergeif Island, mouth of Stikine River (ibid.); Sumdum Village (ibid.); Port Snettisham (ibid.).

=Sorex vagrans mixtus= Hall

Sorex obscurus mixtus Hall, American Nat., 72:462, September 10, 1938.

Type.--Adult male, skin and skull; No. 70376, Mus. Vert. Zool.; obtained on May 4, 1936, by R. A. Cumming, from Vanada, Texada Island, Georgia Strait, British Columbia.

Range.--Known only from the type locality.

Diagnosis.--Size medium; average and extreme measurements of 5 specimens from the type locality are: total length, 111 (108-117); tail, 48 (44-49); hind foot, 12 (12-13) (Hall, 1938:463).

Comparisons.--Color much as in S. v. setosus or S. v. isolatus; palate longer than that of isolatus or setosus; hind foot shorter than either; smaller than S. v. longicauda.

=Sorex vagrans isolatus= Jackson

Sorex obscurus isolatus Jackson, Jour. Washington Acad. Sci., 12:263, June 14, 1922.

Type.--Adult male, skin and skull; No. 177719, U. S. Biol. Surv. Coll.; obtained on May 21, 1911, by Alexander Wetmore from mouth of Millstone Creek, Nanaimo, Vancouver Island, British Columbia.

Range.--Vancouver Island.

Diagnosis.--Size medium; measurements of two from the type locality are: total length, 113, 118; tail, 48, 49; hind foot, 14, 14. Dark in summer and winter, underparts brownish.

Comparisons.--Smaller than S. v. setosus but color much the same; resembles S. v. obscurus in size and cranial characters but darker in all pelages; similar in color to S. v. vancouverensis with which isolatus is sympatric but with longer tail, longer hind feet, broader rostrum and larger teeth. For comparison with S. v. mixtus see account of that subspecies.

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