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Speciation of the Wandering Shrew · James S. Findley — chapter 13 of 21 · ~1,091 words · public domain

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Remarks.--The subspecies longiquus is obviously derived from the neighboring S. v. obscurus and differs from it mainly in size. Some specimens of obscurus from western Montana show evidences of intergradation with S. v. vagrans in possessing a somewhat buffy belly and these are thus more strikingly different from longiquus than are other specimens of obscurus. Many specimens of obscurus from the eastern slope of the Lewis and Clark Range in Montana show the tricolored pattern seen in many specimens of longiquus. The smallest individuals of longiquus are found on the Big Snowy Mountains. Intergradation with obscurus is seen in specimens here referred to S. v. obscurus from the Big Belt Mountains.

Specimens examined.--Total number, 45. MONTANA: Hill Co.: Bearpaw Mts., 5 UM, 2 BS. Phillips Co.: Zortman, 1 BS. Chouteau Co.: type locality, 3 UM; Highwood Mts., 13 BS. Cascade Co.: Neihart, Little Belt Mts., 1 BS. Judith Basin Co.: 3 mi. W Geyser, 4100 ft., 1 KU; Otter Creek, 10 mi. SW Geyser, 1 BS; Dry Wolf Creek, 20 mi. SW Stanford, 1 BS. Buffalo, 13 mi. W Buffalo Canyon, 2 BS. Fergus Co.: Moccasin Mts., 15 mi. NW Hilger, 3 BS; Judith Mts., 17 mi. NE Lewiston, 1 BS; 15 mi. S Heath, N. fork Flat Willow Creek, Big Snowy Mts., 1 BS; Timber Creek, Big Snowy Mts., 1 BS; Crystal Lake, 6000 ft., Big Snowy Mts., 2 UM; Rocky Creek, 5600 ft., Big Snowy Mts., 3 UM; Big Snowy Mts., 3 BS. Meagher Co.: Sheep Creek, 16 mi. N White Sulphur Springs, Little Belt Mts., 1 BS.

Marginal records.--MONTANA: Bearpaw Mts.; Zortman; Big Snowy Mts.; 16 mi. N White Sulphur Springs; Highwood Mts.

=Sorex vagrans neomexicanus= Bailey

Sorex obscurus neomexicanus Bailey, Proc. Biol. Soc. Washington, 26:133, May 21, 1913.

Type.--Adult male, skin and skull; No. 100440, U. S. Biol. Surv. Coll.; obtained on May 29, 1900, by Vernon Bailey, from Cloudcroft, 9000 ft., Otero Co., New Mexico.

Range.--Sacramento and Capitan Mountains of New Mexico.

Diagnosis.--Size medium for the species; average and extreme measurements of 4 topotypes are: total length, 105.2 (103-107); tail, 41.0 (39-42); hind foot, 13.1 (12.5-14). Color near Olive Brown in summer; winter pelage unknown; skull large and relatively broad; teeth relatively large.

Comparisons.--Skull larger than that of S. v. obscurus and relatively somewhat broader; much larger in all cranial dimensions than S. v. monticola.

Remarks.--S. v. neomexicanus is a well-marked subspecies seemingly limited to the mountains of southeastern New Mexico. It is the only species of Sorex thus far recorded from that area.

Specimens examined.--Total number, 12. NEW MEXICO: Otero Co.: SW slope Capitan Mts., 2 BS; 10 mi. NE Cloudcroft, 2 BS; Cloudcroft, 7 BS, 1 UM.

Marginal records.--NEW MEXICO: SW slope Capitan Mts.; 10 mi. NE Cloudcroft; type locality.

=Sorex vagrans monticola= Merriam

Sorex monticolus Merriam, N. Amer. Fauna, 3:43, September 11, 1890.

Sorex vagrans monticola, Merriam, N. Amer. Fauna, 10:69, December 31, 1895.

Sorex melanogenys Hall, Jour. Mamm., 13:260, August 9, 1932, type from Marijilda Canyon, 8600 ft., Graham Mts. [= Pinaleno Mts.] Graham Co., Arizona.

Type.--Adult male, skin and skull; No. 17599/24535, U. S. Biol. Surv. Coll.; obtained on August 28, 1899, by C. Hart Merriam and Vernon Bailey from San Francisco Mtn., 11,500 ft., Coconino Co., Arizona.

Range.--Mountains of western New Mexico, eastern Arizona, and the northern Sierra Madre Occidental of Mexico.

Diagnosis.--Size small for the species; average and extreme measurements of 12 specimens from the White Mountains, Arizona, are: total length, 104.3 (98-112); tail, 41.2 (37-45); hind foot, 12.0 (11-13). Summer pelage between (15'm) Proutts Brown and (15''m) Bister, venter tinged with (15'f) Pale Ochraceous Buff; winter pelage near (17'''k) Olive Brown; skull relatively broad.

Comparisons.--For comparisons with S. v. obscurus and S. v. neomexicanus see accounts of those subspecies. Skull slightly larger and relatively broader than that of S. v. orizabae, and color slightly paler. Differs from S. v. vagrans in: winter pelage grayish (near 17'''k Olive Brown) rather than blackish (17''''k or 17''''m Chaetura Drab or Chaetura Black); summer pelage slightly grayer; skull relatively slightly broader rostrally and interorbitally.

Remarks.--S. v. monticola intergrades gradually with S. v. obscurus to the north and east; indeed the type locality is actually in this area of intergradation. So far as I know, monticola is not in reproductive continuity with any other subspecies of Sorex vagrans. Specimens from southeastern Arizona are the smallest and seem to be the most "typical" in the sense that they are most different from S. v. obscurus. Some specimens from the whole length of the Rocky Mountain chain in the United States have for years been referred to monticola. Some of these, as I have pointed out, belong to S. v. longiquus, and others are intergrades between S. v. obscurus and S. v. vagrans. Since vagrans and monticola resemble one another somewhat, and since topotypes of S. v. monticola actually show the influence of intergradation with obscurus, it is easy to understand how intergrades between obscurus and vagrans could have been assigned to monticola.

Throughout most of its range, S. v. monticola is the only Sorex present. In some places monticola may occur with S. nanus or S. merriami. S. v. monticola occurs with the water shrew in southeastern Arizona.

Specimens examined.--Total number, 80.

ARIZONA: Coconino Co.: San Francisco Mtn., 3 BS, 6 CMNH. Apache Co.: Spruce Creek, Tunitcha Mts., 7 BS; Springerville, 1 BS; North Fork White River, White Mts., 12 SD; White River, Horseshoe Cienega, 8300 ft., White Mts., 5 BS; Mt. Thomas, 9500 to 11,000 ft., White Mts., 12 BS; Little Colorado River, White Mts., 4 BS; near head Burro Creek, 9000 ft., White Mts., 1 BS. Graham Co.: Graham Mts., 9200 ft., 2 BS. Greenlee Co.: Prieto Plateau, 9000 ft., S. end Blue Range, 1 BS. Pima Co.: Summerhaven, 7500 ft., Santa Catalina Mts., 3 BS, 1 SD. Cochise Co.: Fly Park, Chiricahua Mts., 4 BS; Rustler Park, Chiricahua Mts., 1 SD; Long Park, Chiricahua Mts., 1 UM; Huachuca Mts., 1 BS. Santa Cruz Co.: Stone Cabin Canyon, 8500 ft., Santa Rita Mts., 1 BS.

NEW MEXICO: San Juan Co.: Chusca Mts., 1 BS. Catron Co.: Mogollon Mts., 3 BS; 10 mi. E Mogollon, 1 KU. Socorro Co.: Copper Canyon, Magdalena Mts., 3 BS. Sierra Co.: Mimbres Mts., near Kingston, 1 BS.

CHIHUAHUA: Sierra Madre, near Guadalupe y Calvo, 5 BS.

Marginal records.--ARIZONA: Tunitcha Mts. NEW MEXICO: Chusca Mts.; Copper Canyon, Magdalena Mts.; Mimbres Mts., near Kingston. CHIHUAHUA: Guadalupe y Calvo. ARIZONA: Huachuca Mts.; Santa Catalina Mts.; White River, Horseshoe Cienega, 8300 ft., White Mts.; San Francisco Mtn.

=Sorex vagrans orizabae= Merriam

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