Part 2

Dentition larger and heavier than that in rotundus, but otherwise differing only slightly from it; upper incisor less concave on cutting surface (see Figs. 3-4); premolar and molar slightly less bladelike, with heavier roots.

The peculiar shape of the incisor of stocki is shared to some extent with Diaemus youngi, a Recent South American desmodontid. However, stocki does not otherwise resemble D. youngi, differing from it as follows: Skull larger and heavier; interorbital constriction much narrower; zygomatic arches less strongly bowed; skull less compact, more elongate; braincase and rostrum relatively much narrower in relation to greatest length of skull. Furthermore, specimens of stocki show no trace of the minute M2 attributed to youngi by de la Torre (Proc. Biol. Soc. Washington, 69: 191, 1956). For cranial measurements of youngi see Sanborn (Jour. Mamm., 30: 283, 1949).

Remarks.--The essential differences between D. stocki and D. rotundus are in size and proportion. I do not doubt that the two species are closely related; possibly stocki is ancestral to rotundus.

The species is named in honor of the late Professor Chester Stock, under whose direction the fossil materials from San Josecito Cave were obtained, and who, at the time of his death, was studying the mammalian fauna from the cave.

Eptesicus cf. grandis (Brown)

Referred material.--One rostrum, with P4-M3 on the right side and P4 only on the left, LACM (CIT) 2990.

Remarks.--This specimen is referred provisionally to E. grandis. The dentition is larger and heavier, and the ridges and depressions on the dorsal surface of the rostrum are more pronounced than in Recent E. fuscus. The P4-M3 length is 6.1 (approximately 6.1 in the holotype of grandis, less in fuscus); least interorbital constriction, 4.2 (4.3 in the holotype of grandis, more in fuscus); breadth of rostrum between infraorbital canals, 6.4; breadth across P4, 7.3.

TABLE 1.--Cranial measurements of two species of Desmodus.

---------------------+-------+-------+-------+-------+-------+-------- Catalogue number | G o | C l | Z b | B b | L c | B f or number of | r f | o e | y r | r r | e o | r o specimens averaged | e | n n | g e | e a | a n | e r | a s | d g | o a | a i | s s | a a | t k | y t | m d | d n | t t | d m | e u | l h | a t | t c | r | t e | s l | o | t h | h a | i i | h n | t l | b | i | s | n c | | | a | c | o e | t t | o m | l | s | | f | e i | f a | e | a | | | r o | g | n | l | | | o n | n | g | | | | r | u | t | | | | b | m | h | | | | i | | | | | | t | | | | | | a | | | | | | l | ---------------------+-------+-------+-------+-------+-------+--------

Desmodus rotundus murinus, La Mula, 13 mi. N Jaumave, Tamaulipas

10 (3 male, Ave. | 24.3 | 21.4 | 12.0 | 12.1 | 5.5 | 5.2 7 female) Max. | 24.9 | 22.0 | 12.5 | 12.5 | 5.6 | 5.3 Min. | 23.9 | 21.0 | 11.7 | 11.9 | 5.3 | 5.1

Desmodus stocki, San Josecito Cave, Nuevo León

2946 | 27.3 | 24.5 | | 14.2 | 6.1 | 5.8 2947 | | | | 13.6 | | 5.7 2948 | | 24.3 | | 13.9 | 6.2 | 5.3 2949 | | 24.7 | | 13.9 | 6.1 | 5.5 2950 | | | 14.1 | 13.5 | | 5.7 3127 | | | | 13.5 | 6.0 | 5.7 3128 | 26.5 | | | 13.5 | 6.2 | 5.5 3129 (type) | 28.2 | 24.5 | | 13.7 | 5.9 | 5.7 3940 | 27.4 | 24.4 | | 13.8 | 6.2 | 3941 | | 24.6 | 14.0 | 13.7 | 6.0 | 5.6 ---------------------+-------+-------+-------+-------+-------+--------

Brown (1908:174) originally named grandis as a subspecies of fuscus. Gidley and Gazin (1938:11) considered it a distinct species. Whether grandis is only a subspecies of E. fuscus or a separate species, grandis is closely related to fuscus, and probably is ancestral to it.

Lasiurus cinereus (Palisot de Beauvois)

Referred material.--One cranium, lacking basioccipital, tympanic and mastoid regions, and most of the dentition, having only M3 on the right side and M2-M3 on the left, LACM (CIT) 3160.

Remarks.--The cranium of No. 3160 is inseparable from those of 10 spring-taken specimens of L. c. cinereus from the San Gabriel Mts., Los Angeles Co., California (KU 49727, 49729-37). Measurements of No. 3160, followed by the average and extremes (in parentheses) of the Californian series, are: Condylobasal length, 16.1, 16.5 (15.9-17.2); zygomatic breadth, 12.3, 12.4 (12.0-12.7); least interorbital constriction, 5.2, 5.4 (5.2-5.6); breadth of braincase, 8.7, 9.0 (8.5-9.3); length of palate not including terminal spine, 5.1, 5.3 (4.8-5.9). The teeth of the San Josecito specimen are comparatively unworn. A label with the skull bears the notation "talus" in parentheses, which, in so far as I am able to determine, indicates surface talus inside the cave. Therefore, the specimen in question may be of Recent origin.

It is perhaps worthy of note that Lasiurus cinereus is primarily a tree-dwelling bat, although a few Recent specimens have been reported from caves (see Beer, 1954:116).

Corynorhinus tetralophodon Handley

A single cranium of a Corynorhinus LACM (CIT) 2989 was included in the original materials sent to Kansas by Professor Stock. Subsequently, this specimen was loaned to Charles O. Handley, Jr., who described it as a new species, C. tetralophodon. The latter is said to differ from all other plecotine bats by the retention of a well-developed fourth commissure (ridge extending posteroexternally from metacone) on the M3 (Handley, 1955:48).

LITERATURE CITED

BEER, J. R.

1954. A record of the hoary bat from a cave. Jour. Mamm., 35:116, February 10.

BROWN, B.

1908. The Conard Fissure, a Pleistocene bone deposit in northern Arkansas: with description of two new genera and twenty new species and subspecies of mammals. Mem. Amer. Mus. Nat., 9:155-208, pls. 14-25.

CUSHING, J. E., JR.

1945. Quaternary rodents and lagomorphs of San Josecito Cave, Nuevo Leon, Mexico. Jour. Mamm., 26:182-185, July 19.

FINDLEY, J. S.