=42.= Other structures besides the diaphragm assist in pumping air in and out of the lungs. By the action of the diaphragm the chest is alternately lengthened and shortened. But if you watch anyone, and especially a woman, breathing, you will notice that with every breath the chest rises and falls; the front of the chest, the sternum, as you have learnt to call it, comes forward and goes back; and a little attention will convince you that it comes forward during inspiration, i.e. while the diaphragm is descending, and falls back during expiration. But this coming forward of the sternum means a widening of the chest from back to front, and the falling back of the sternum means a corresponding narrowing. So that while the chest is being lengthened by the descent of the diaphragm, it is also being widened by the coming forward of the sternum. In inspiration the lungs are expanded not only downwards, by the movement of the diaphragm, but also outwards, by the movement of the walls of the chest.
What thrusts forward the sternum? If you were to watch closely the sides of the chest of a very thin person, you would be able to notice that at every breathing in, at every inspiration, the ribs are pulled up a little way. Now, each rib is connected with the backbone behind by a joint, and is firmly fastened to the sternum in front by cartilage (see Frontispiece). If you were to fasten a piece of string to the middle of one of the ribs and to pull it, you would find you were working on a lever, with the fulcrum at the backbone, with the weight acting at the sternum, and the power at the point where your string was tied. Every time you pulled the string the rib would move on its fulcrum at the backbone, in such a way that the front end of the rib would rise up, and the sternum would be thrust out a little. When you left off pulling, the sternum, which in being thrust forward had been put on the stretch, would sink back, and the rib would fall down to its previous position.
a. The bony rib; b, the cartilage; c, the junction of bone and cartilage; d, unossified; e, ossified, portions of the sternum. A. External intercostal muscle. B. Internal intercostal muscle. In the middle interspace, the external intercostal has been removed to show the internal intercostal beneath it. ]
Between the ribs are certain muscles called =intercostal muscles= (Fig. 14). The exact action of these you will learn at some future time. Meanwhile it will be enough to say that they act like the piece of string we are speaking of. =When they contract, they pull up the ribs and thrust out the sternum; when they leave off contracting, the ribs and sternum fall back to their previous position.=
There are many other muscles which help in breathing, especially in hard or deep breathing, but it will be sufficient for you to remember that in ordinary breathing there are two chief movements taking place exactly at the same time, by means of which air is drawn into the chest, both movements being caused by the contraction of muscles. First, the diaphragm contracts and flattens itself, making the chest deeper or longer; secondly, at the same time the ribs are raised and the sternum thrust out by the contraction of the intercostal muscles, making the chest wider. But as the chest becomes wider and longer, the lungs become wider and longer too. In order to fill up the extra room thus made in the lungs, air enters into them through the windpipe. This is =inspiration=. But soon the diaphragm and the intercostal muscles cease to contract; the diaphragm returns to its arched condition, the ribs sink down, the sternum falls back, and the extra air rushes back again out of the lungs through the windpipe. This is =expiration=. An inspiration and an expiration make up a whole breath; and thus we breathe some seventeen times in every minute of our lives.
=43.= But what makes the diaphragm and intercostal muscles contract and rest in so beautifully regular a fashion? The biceps of the arm, we saw, was made to contract by our will. It is not our will, however, which makes us breathe. We breathe often without knowing it; we breathe in our sleep when our will is dead; we breathe whether we will or no, because we cannot help it. We can quicken our breathing, we can take a short or deep breath as we please, we can change our breathing by the force of our will; but the breathing itself goes on without, and in spite of, our will. It is =an involuntary act.=
Though breathing is not an effort of the will, it is an effort of the brain; an effort, too, of one particular part of the brain, that part where the brain joins on to the spinal cord. Nerves run from the diaphragm and the intercostal and other muscles through the spinal cord, to this part of the brain. And seventeen times a minute a message comes down along these nerves, from the brain, bidding them contract; they obey, and you breathe. Why and how that message comes, you will learn at some future time. When your head is cut off, or when that part of the brain which joins on to the spinal cord is injured by accident or made powerless by disease, the message ceases to be sent, and you cease to breathe.
=44.= At every breath, then, a certain quantity of air goes in and out of the chest; but only a small quantity. =You must not think the lungs are quite emptied and quite filled at each breath.= On the contrary, you only take in each time a mere handful of air, which reaches about as far as the large branches of the windpipe, and does not itself go into the air-cells at all. This is often called =tidal= air; and the rest of the air in the lungs, which does not move, is often called the =stationary= air (see Fig. 13).
How then does the carbonic acid at the bottom of the lungs get out? How do the capillaries in the air-cells get their fresh oxygen?
The stationary air mingles with the tidal air at every breath. If you want to ventilate a room, you are not obliged to take a pair of bellows and drive out every bit of the old air in the room, and supply its place with new air: it will be enough if you open a window or a door and let in a draught of pure air across one corner, say, of the room. That current of pure air flowing across the corner will mingle with all the rest of the air until the whole air in the room becomes pure; and the mingling will take place very quickly. So it is in the lungs. The tidal air comes in with each inspiration as pure air from without; but before it comes out at the next expiration it gives up some of its oxygen to the stationary air, and robs the stationary air of some of its carbonic acid. For each breath of tidal air the stationary air is so much the better, having lost some of its carbonic acid and gained some fresh oxygen. The tidal air rapidly purifies the stationary air, and the stationary air purifies the blood.
Thus it comes to pass that the tidal air, which at each pull of the diaphragm and push of the sternum goes into the chest as pure air with twenty-one parts oxygen to seventy-nine parts nitrogen in every hundred parts, comes out, when the diaphragm goes up and the sternum falls back, as impure air with only sixteen parts oxygen, but with five parts carbonic acid to seventy-nine of nitrogen. That lost oxygen is carried through the stationary air to the blood in the capillaries, and the gained carbonic acid came through the stationary air from the blood in the capillaries. So each breath helps to purify the blood, and the pumping of air in and out of the chest changes the impure, hurtful, venous, to pure, refreshing, arterial blood; the blood breathes air in the lungs, that all the body may in turn breathe blood.
HOW THE BLOOD IS CHANGED BY FOOD: DIGESTION. § VII.
=45.= The blood is not only purified by air, it is also renewed and made good by food. The food we eat becomes blood. But our food, though frequently moist, is for the most part solid. We cut it into small pieces on the plate, and with our teeth we crush and tear it into still smaller morsels in our mouth. Still, however well chewed, a great deal of it, most of it in fact, is swallowed solid. In order to become blood it must first be dissolved. It is dissolved in the alimentary canal, and we call the dissolving =digestion=. Let us see how digestion is carried on.
Your skin, though sometimes quite moist with perspiration, is as frequently quite dry. The inside of your mouth is always moist--very frequently quite filled with fluid; and even when you speak of it as being dry, it is still very moist. Why is this? The inside of your mouth is also very much redder than your skin. The redness and the moisture go together.
In speaking of the capillaries, I said that almost all parts of the body were completely riddled with them, but =not quite all=. A certain part of the skin, for instance, has no capillaries or blood-vessels at all. You know that where your skin is thick, you can shave off pieces of skin without “fetching blood;” if your
a, horny epidermis; b, softer layer; c, dermis; d, lowermost vertical layer of epidermic cells; e, cells lining the sweat duct continuous with epidermic cells; h, corkscrew canal of sweat duct. To the right of the sweat duct the dermis is raised into a papilla, in which the small artery, f, breaks up into capillaries, ultimately forming the veins, g. ]
knife were very sharp and you very skilful, you might do the same in every part of your skin. If you were to put some of the skin you had thus cut off under the microscope, you would find that it was made up of little scales. And if you were to take a very thin upright slice running through the whole thickness of the skin, and examine that under a high power of the microscope, you would find that the skin was made up of two quite different parts or layers, as shown in Fig. 15. The upper layer, a, b, is nothing but a mass of little bodies packed closely together. At the top they are pressed flat into scales, but lower down they are round or oval, and at the same time soft. They are called =cells=. As you advance in your study of Physiology you will hear more and more about cells. This layer of cells, either soft and round, or flattened and dried into scales, is called the =epidermis=. No blood-vessel is ever found in the epidermis, and hence, when you cut it, it never bleeds. As long as you live it is always growing. The top scales are always being rubbed off. Whenever you wash your hands, especially with soap, you wash off some of the top scales; and you would soon wash your skin away, were it not that new round cells are always being formed at the bottom of the epidermis, along the line at d (Fig. 15), and always moving up to the top, where they become dried into scales. Thus the skin, or more strictly the epidermis, is always being renewed. Sometimes, as after scarlet fever, the new skin grows quickly, and the old skin comes away in great flakes or patches.
The lower layer below the epidermis is what is called the =dermis=, or =true skin=. This is full of capillaries and blood-vessels, and when the knife or razor gets down to this, you bleed. It is not made up of cells like the epidermis, but of that fibrous substance which you early learnt to call connective tissue (see p. 9). Its top is rarely level, but generally raised into little hillocks, called =papillæ=, as in the figure; the epidermis forming a thick cap over each papillæ, and filling up the hollows between them. Most of the papillæ are full of blood-vessels.
Now, then, I think you will understand why your skin is not red, but flesh-coloured, and why it is generally dry. The true skin under the epidermis is always moist, because of the blood-vessels there; the waste and fluid parts of the blood pass readily through the walls of the capillaries, as you have learnt, by osmosis, and so keep everything round them moist. But this moisture is not enough to soak through the thick coating of epidermis, and so the top part of the epidermis remains dry and scaly.
The true skin underneath the epidermis is always red; you know that if you shave off the surface of your skin anywhere, it gets redder and redder the deeper you go down, even though you do not fetch blood. It is red because of the immense number of capillaries, all full of red blood, which are crowded into it. When you look at these capillaries through a great thickness of epidermis, the redness is partly hidden from you, as when you put a sheet of thin white paper over a red cloth, and the skin seems pink or flesh-coloured; and where the epidermis is very thick, as at the heel, the skin is not even pink, but white or yellow, more or less dirty according to circumstances.
=46.= But if the moist true skin is thus everywhere covered by a thick coat of epidermis, which keeps the moisture in, how is it that the skin is nevertheless sometimes quite moist, as when we perspire?
a, the coil; b, the duct; c, network of capillaries, inside which the duct gland lies.]
If you look at Fig. 15, you will see that the epidermis is at one point pierced by a canal (h) running right through it. You will notice that this canal is not closed at the bottom of the epidermis, but runs right into the dermis or true skin, where the canal becomes a tube, with just one layer (e) of cells, like the cells of the epidermis, for its walls. There is no room in Fig. 15 to show what becomes of this tube, but it runs some way down under the skin all among the blood-vessels, and then twisting itself up into a knot, ends blindly, as is shown in Fig. 16, where b is a continuation on a smaller scale of the same tube which is seen in Fig. 15. This knot is covered by a close network of capillaries, which at c are supposed to be unravelled and taken away from the knotted tube in order to show them. The capillaries, you will understand, though inside the knot, are always outside the tube. If you were to drop a very diminutive marble in at h (Fig. 15), it would rattle down the corkscrew passage through the thick epidermis, shoot down the straight tube b (Fig. 16), and roll through the knot a, until it came to rest at the blind end of the tube. Along its whole course it would touch nothing but cells, like the cells of the epidermis, a single layer of which forms the walls of the tube where it runs below the epidermis. If it got lodged at h (Fig. 15), or got lodged in the knot at a (Fig. 16), it would in both cases be touching epidermic cells. But there would be this great difference. At h it would be ever so far removed from any blood capillary; at a it would only have to make its way through a thin layer of single cells, and it would be touching a capillary directly. At h it might remain dry for some time; at a it would get wet directly, for there is nothing to prevent the fluid parts of the blood oozing out through the thin wall of the capillaries, and so through the thin wall of the tube into the canal of the tube, on to the marble.
In fact, the inside of the knot is always moist and filled with fluid. When the capillaries round the knot get over-full of blood, as they often do, a great deal of colourless watery fluid passes from them into the tube. The tube gets full, the fluid wells up right into the corkscrew portion in the thickness of the epidermis, and at last overflows at the mouth of the tube over the skin. We call this fluid sweat or =perspiration=. We call the tube with its knotted end =a gland=; and we call the act by which the colourless fluid passes out of the blood capillaries into the canal of the tube, =secretion=. We speak of the =sweat gland secreting sweat out of the blood brought by the capillaries which are wrapped round the gland=.
=47.= Now we can understand why the inside of the mouth is red and moist. The mouth has a skin just like the skin of the hand. There is an outside epidermis, made up of cells and free from capillaries, and beneath that a dermis or true skin crowded with capillaries. Only the epidermis of the mouth is ever so much thinner than that of the hand. The red capillaries easily shine through it, and their moisture can make its way through. Hence the mouth is red and moist. Besides there are many glands in it, something like the sweat gland, but differing in shape; these especially help to keep it moist.
Because it is always red and moist and soft, the skin of the inside of the mouth is generally not called a skin at all, but =mucous membrane=, and the upper layer is not called epidermis, but =epithelium=. You will remember, however, that a mucous membrane is in reality a skin in which the epidermis is thin and soft, and is called epithelium.
The mouth is the beginning of the alimentary canal. Throughout its whole length the alimentary canal is lined by a skin or mucous membrane like that of the mouth, only over the greater part of it the epithelium is still thinner than in the mouth, and indeed is made up of a single layer only of cells. The whole of the inside of the canal is therefore red and moist, and whatever lies in the canal is separated by a very thin partition only from the blood in the capillaries, which are found in immense numbers in the walls of the canal. The alimentary canal is, as you know, a long tube, wide at the stomach but narrow elsewhere. In all parts of its length the tube is made up of mucous membrane on the inside, and on the outside of muscles, differing somewhat from the muscles of the body and of the heart, but having the same power of contracting, and by contracting of squeezing the contents of the tube, just as the muscles of the heart squeeze the blood in its cavities. The muscles, and especially the mucous membrane, are crowded with blood-vessels.
Though the epithelium of the mucous membrane is very thin, the mucous membrane itself is thick, in some places quite as thick as the skin of the body. This thickness is caused by its being =crowded with glands=. In the skin the sweat glands are generally some little distance apart, but in the mucous membrane of the stomach and of the intestines they are packed so close together, that the membrane seems to be wholly made up of glands.
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