Diagnosis.--Bill short, flat, somewhat obtuse, minutely notched near tip of each maxilla, flared at base; gape wide and deeply cleft; culmen convex; nasal fossa broad, exposed, or filled with short, erect or antrorse, close-set velvety feathers; nostril narrowly elliptical; rictal vibrissae long, short, or absent; lacrimal bone free, articulating at two points; wings long and pointed, or short and rounded; primaries ten, tenth reduced in some species; tail short, narrow, even, two thirds or less length of wing, or much longer and forked or rounded; feet weak (except in Dulus and Phainoptila); tarsus generally shorter than middle toe and claw, distinctly scutellate with five or six divisions, the lateral plate subdivided (except in Phainoptila); lateral toes of nearly equal length; hallux approximately as long as inner lateral toe, or shorter; basal phalanx of middle toe more or less united to that of outer and inner toes; body stout; head generally conspicuously crested; plumage soft, smooth and silky (except in Dulus); eggs spotted; nest in trees; three subfamilies, five genera, eight species.
Subfamily Ptilogonatinae
Diagnosis.--Rictus with conspicuous bristles; nasal fossa almost entirely exposed; tail long and rounded, graduated, or square; caudal muscles and pygostyle well developed; wings rounded and short, first primary a half to a third as long as second; second primary shorter than third; humerus long, with small external condyle; plumage soft and silky, less so in Phainoptila; sexes dissimilar, young like adult female; three genera, four species.
Genus =Phainoptila= Salvin
Phainoptila Salvin, Proc. Zool. Soc. London, 1877:367, April 17, 1877. Type Phainoptila melanoxantha Salvin.
Diagnosis.--Without crest; tarsus longer than middle toe and claw, and booted or very slightly reticulate; tail shorter than wing, rounded; nostril exposed, ovate; rictal bristles distinct; first primary well developed; plumage normal, bill flared slightly at base.
Range.--Costa Rica and Panama.
=Phainoptila melanoxantha melanoxantha= Salvin
Phainoptila
Phainoptila melanoxantha melanoxantha Salvin, Proc. Zool. Soc. London, 1877:367; April 17, 1877.
Diagnosis.--Coloration of adult males: Pileum, hindneck, back, scapulars, and upper tail coverts Black (capitalized color terms after Ridgway, Color Standards and Color Nomenclature, Washington, D. C., 1912), with Bluish Gray-Green gloss; rump Lemon Yellow tinged with Olive; lower breast and abdomen Gull Gray or Slate Gray; sides and flanks clear Lemon Yellow; lower chest, upper breast, and under tail coverts Yellowish Olive-Green, extending to patch on sides and flanks of same color; bill and feet Black or Blackish Brown. Coloration of adult females: Most of upper parts Olive-Green, with Yellowish Olive on rump; thighs Olive-Gray, as are sides of head; rest of coloration as in male. Coloration of young: As in adult female, but duller throughout.
Measurements.--Wing 99.0, tail 88.5, culmen 15.2, tarsus 28.4.
Range.--Highlands of Costa Rica and extreme western Panama (Volcan de Chiriqui).
=Phainoptila melanoxantha minor= Griscom
Phainoptila
Phainoptila melanoxantha minor Griscom, Amer. Mus. Novitates, 141:7, 1924.
Diagnosis.--Coloration as in P. m. melanoxantha, but female with hindneck more extensively gray and of slightly darker shade; rump, upper tail coverts, and edgings to tail feathers slightly greener, less yellow; average size smaller than in P. m. melanoxantha.
Range.--Highlands of westeran Panama (Cerro Flores and eastern Chiriqui).
Genus =Ptilogonys= Swainson
Ptilogonys Swainson, Cat. Bullock's Mex. Mus., App. 4, 1824. Type Ptilogonys cinereus Swainson.
Diagnosis.--Tail much longer than wing, even or graduated; head with bushy crest; nostril large, rounded and fully exposed, bordered by membrane; rictal bristles well developed; tarsus shorter than middle toe with claw; plumage soft, blended.
Range.--Southwestern United States to Costa Rica.
=Ptilogonys cinereus cinereus= Swainson
Ashy Ptilogonys
Ptilogonys cinereus cinereus Swainson, Cat. Bullock's Mex. Mus., App. 4, 1824.
Diagnosis.--Coloration of adult male: Frontals, supralorals, malars, and chin White; orbital ring White; auriculars and nape grayish brown; rest of head smoke gray; back, scapulars, wing coverts, rump, and upper tail coverts plain Bluish Black; rectrices (except middle pair) with large patch of White midway between base and tip, rest plain Bluish Black; chest, breast, and anterior parts of sides plain Bluish Gray-Green, much lighter than back, and fading into paler Gray on throat; abdomen and thighs White; flanks and posterior part of sides Olive-Yellow or Yellowish Olive; under tail coverts Lemon Yellow; bill, legs and feet Black. Coloration of adult females: Head plain Smoke Gray, passing into White on frontals, malars, and chin; back, scapulars, wing coverts, and rump Hair Brown; upper tail coverts Dark Gull Gray; remiges and rectrices Black with faint Dusky Green gloss, edged with Gull Gray; chest Dark Grayish Brown lightening to Wood Brown on sides and flanks; abdomen White; under tail coverts Yellow Ocher. Coloration of young: As in adult female, but paler throughout.
Measurements.--In adult male, wing 94.0, and tail 104.2; in adult female, wing 93.3, and tail 94.8; both sexes, culmen 11.1, and tarsus 18.7.
Range.--Mountainous districts of central and southern Mexico, in states of Durango, Zacatecas, Hidalgo, Mexico, Oaxaca, Colima, Morelos, Veracruz, San Luis Potosi, Guerrero and Michoacan.
Phylogeny of the Waxwings and Allied Birds · The Wunder Library — complete classics, free to read, with narration.