Part 8

Again, the geography of the Globe changed. New lands arose from the sea, and old lands partially or wholly sank. The German Ocean, and part of Western Europe, of our maps, were a great group of islands. The Oolitic formation was deposited. The general character of the organization of this period differed little from that of the Lias. New forms of plants, such as Cycadeae, were abundant, with, considerable numbers of Corals, Encrinites, Sea-urchins and Mollusks. Macrurous Crustacea, much like those of our times (but essentially different in species), inhabited the sea, and some Beetles and Flies represented the Insects of the land. The Fishes and Marine Reptiles were pretty much the same with those of the Lias, though they received some important additions.

It is, however, among the terrestrial Vertebrata that we must look for the characteristic organisms of this age. And these are, still, Reptiles. The huge Megalosaurus, with a body as big as an elephant's, stood high on his legs, and stretched open a pair of gaping jaws, set with jagged teeth. The Pterodactyles flew about,--carnivorous lizards, with the body and wings of bats, except that the membrane was stretched upon the enormously developed little finger;--creatures, perhaps, the most unlike to anything familiar to us, of all fossil forms. And, in the marshy margins of the great river valley which formed the Wealden of our South-eastern districts, the giant Iguanodon, and his fellow, the Hylaeosaurus, waged their peaceful warfare on the succulent plants that became their unresisting prey.

Pterodactylus crassirostris, and P. brevirostris.]

The circle of animal life was completed in this epoch, thus far, that every class was represented by some one or more of its constituent species. No fossil skeletons of Birds have, indeed, been found so low as the Oolite, but numerous foot-prints of some of the Grallatores are found in a sandstone of this period; and in the Stonesfield slate, which is contemporary with it, a genus of Mammalia has been discovered,--a small Marsupial, allied to the Opossums of America.

The duration of the Oolitic period must have been considerable. "The lias sea-bottom was succeeded first by a sandy, and then by a calcareous deposit, and the animals were modified accordingly." The deposit of carbonate of lime, which took place under circumstances that caused it to attract around its nodules the organic particles, whence the name oolite (egg-stone) is derived, was not continuous, but repeated at intervals. The shells of Mollusks were developed in great abundance, and accumulations of these formed thick bands, which consolidated into layers of shell-limestone. Three hundred feet of strata, largely composed of organic remains, were formed before the clay was deposited which made the Stonesfield and contemporaneous slates.

Once more the dry land sank, probably by slow successive subsidences, and the sea flowed many fathoms deep above the great European archipelago. And upon its quiet bottom settled down, first a few sandy and clayey beds, and then the great layer of the Chalk.

Creatures of very minute size and low grades of organization were now playing a very important part. A large portion of the lime that was deposited, in the form of a pure carbonate, was doubtless supplied by the Coral structures, which, were exceedingly numerous; the polypidoms being gnawed down by strong-jawed fishes that fed upon the Zoophytes. Foraminifera also were abundant, and contributed to the supply.

Nodules of flint exist in the Chalk, sometimes scattered, sometimes arranged in bands. Two sources are indicated for this substance. One is Sponge, the most common kinds of which are composed of skeletons of siliceous spicula; and these can be discerned with the microscope in the interior of the chalk-flints. But millions upon millions of Infusoria swam through the waters, and many of these were encased in siliceous loricae, while the rocks and sea-weeds were fringed with as incalculably numerous examples of siliceous Diatomaceae, whose elegant forms are recognisable without difficulty throughout the Chalk. The inconceivable abundance of these forms may be illustrated by the often-cited fact, that whole strata of solid rock appear to be so exclusively composed of their solid remains, that a cube of one-tenth of an inch is computed by Ehrenberg to contain five hundred millions of individuals.

The increase of these organisms is very rapid, and their duration proportionately short; but allowing for this, what period would elapse before the successive generations of entities, of which forty-one thousand millions are required to make a cubic inch, would have accumulated into solid strata fourteen feet in thickness?

Without pausing to examine the whole Cretaceous fauna, we may observe that the Mollusca with chambered shells--the Ammonites and their allies--were developed in singular variety and profusion during this period, after which they suddenly disappeared from the ocean. The Fishes present little that is remarkable; of Birds, few, and of Mammals, no remains exist; and the Reptiles, while not absolutely extinct, are few and rare. One great marine form, however, the Mosasaurus, was added to their number.

At length the sea ceased to deposit chalk, and its bed appears to have been slowly elevated, until all the animals that had inhabited the waters of that formation were destroyed; so that their race and generation perished. The grand epoch of Secondary Formations was closed.

It was followed by an extensive disruption of the then existing strata, and by changes and modifications so great as to alter the whole face of nature. "It would appear that a long period of time elapsed before newer beds were thrown down, since the chalky mud not only had time to harden into chalk, but the surface of the chalk itself was much rubbed and worn." During this protracted period, eruptions of molten rock occurred of enormous extent, producing the Basaltic formation which covers the Chalk in the north of Ireland, and in some of the Hebrides. In the south of Europe the Pyrenees were elevated, and the Apennines and Carpathians were pushed to a greater altitude than before, if they were not then formed. The Alps and the Caucasus also experienced a series of upward movements, continuing through a considerable range of the Tertiary epoch.

The rich collections of vegetable remains--chiefly fruits and seeds--that have been made from the London Clay, show that the earliest land of this period was clothed with a great abundance and variety of plants; and these are of such alliances as would now require a tropical climate. Many species of Palms, Screw-pines, Gourds, Piperaceae, Mimoseae, and other Leguminosae, Malvaceae, and Coniferae, dropped their woody pods and fruits where now these pages are written; and the animals manifest no less interesting an approximation to existing forms than the plants. The Zoophytes, the Echinoderms, the Foraminifera, the Worms, the Crustacea, the Mollusca, the Fishes and the Reptiles of the Eocene beds, exhibit a great preponderance of agreement with those that now exist, so far as genus is concerned, though the species are still almost wholly distinct. The approximation is particularly marked in the Molluscous sub-kingdom, by the almost entire disappearance of the hitherto swarming Brachiopod and Cephalopod forms, and the progressive substitution for them of the Conchifera and Gastropoda, which had, however, throughout the Secondary epoch, been gradually coming forward to their present predominance in nature.

Among the Fishes, the Placoid type was diminished in number; and those that were produced were mostly Sharks and Rays, of modern genera; but the chief difference was the paucity of those mailed forms (Ganoids), which were so abundant during the Oolitic period. On the other hand, the Ctenoid and Cycloid forms, which had begun to make their appearance in small numbers in the Chalk, are well represented. In both this deficiency and this plenitude, there is a very decided approach to existing conditions; for the Ganoids are almost unknown with us, while the last-named two orders are abundant. Representatives of our Perches, Maigres, Mackerels, Blennies, Herrings, and Cods, were numerous; distinct, however, from the present species. But not a single member of the great Salmon family was yet introduced.

The great Saurian Reptiles had entirely disappeared, and were quite unrepresented in the tertiary beds, except by a Crocodile or two, and a small Lizard. Turtles were, however, numerous, both of the marine and lacustrine kinds; and there is an interesting stranger, in the form of a large Serpent, allied to our Pythons, some twenty feet in length.

Birds and Mammals began now to assume their place on the land. The London Clay presents us with a little Vulture; and the Paris basin contains remnants of species representing the Raptores, the Rasores, the Grallatores, and the Natatores.

The Quadrupeds came in in some force; not developed from the lowest to the highest scale of organization; for the Monkey and the Bat occur in sands, certainly not later, if not earlier, than the London Clay, contemporaneously with the Racoon, and before the existence of any Rodent or Cetacean. Some Carnivora, as the Wolf and the Fox, roamed the woods, but the character of the epoch was given by the Pachyderms.

These, however, were not the massive colossi that browse in the African or Indian jungles of our days; no Elephant, no Rhinoceros, no Hippopotamus was as yet formed. But several kinds of Tapir wallowed in the morasses; and a goodly number of largish beasts, whose affinities were with the Pachydermata, while their analogies were with the Ruminantia, served as substitutes for the latter order, which was wholly wanting. These interesting quadrupeds, forming the genus Anoplotherium, were remarkable for two peculiarities,--their feet were two-toed, and their teeth were ranged in a continuous series, without any interval between the incisors and the molars. They varied in size from that of an ass to that of a hare.

The physical conditions of our earth, when it was tenanted by these creatures, is thus described:--"All the great plains of Europe, and the districts through which the principal rivers now run, were then submerged; in all probability, the land chiefly extended in a westerly direction, far out into the Atlantic, possibly trending to the south, and connecting the western shores of England with the volcanic islands off the west coast of Africa. The great mountain chains of Europe, the Pyrenees, the Alps, the Apennines, the mountains of Greece, the mountains of Bohemia, and the Carpathians, existed then only as chains of islands in an open sea. Elevatory movements, having an east and west direction, had, however, already commenced, and were producing important results, laying bare the Wealden district in the south-east of England. The southern and central European district, and parts of western Asia, were the recipients of calcareous deposits (chiefly the skeletons of Foraminifera), forming the Apennine limestone; while numerous islands were gradually lifted above the sea, and fragments of disturbed and fractured rock were washed upon the neighbouring shallows or coast-lines, forming beds of gravel covering the Chalk. The beds of Nummulites and Miliolites, contemporaneous with those containing the Sheppey plants and the Paris quadrupeds, seem to indicate a deep sea at no great distance from shore, and render it probable that there were frequent alternations of elevation and depression, perhaps the result of disturbances acting in the direction already alluded to.

"The shores of the islands and main land were, however, occasionally low and swampy, rivers bringing down mud in what is now the south-east of England, and the neighbourhood of Brussels, but depositing extensive calcareous beds near Paris. Deep inlets of the sea, estuaries, and the shifting mouths of a river, were also affected by numerous alterations of level not sufficient to destroy, but powerful enough to modify, the animal and vegetable species then existing; and these movements were continued for a long time."

After the elevation of the mountain summits of Europe above the sea, and while the same causes were still in operation, deposits were being made in the narrow intervening seas of the Archipelago, such as the present south of France, the valleys of the Rhine and Danube, the eastern districts of England and Portugal. These deposits were partly marine and partly lacustrine; the former consisting largely of loose sands, mingled with shells and gravel. In Switzerland is a thick mass of conglomerate; and in the district around Mayence, there is a series of fresh-water limestones, and sandstones charged with organic remains.

The changes which took place during this comparatively recent epoch were not sudden, but gradual; the results of operations which were probably going on without intermission, and perhaps have not yet ceased. The land was more and more upheaved, till at length, what had been an archipelago of islands became a continent, and Europe assumed the form which it bears on our maps.

The most interesting addition to the natural history of the Miocene, or Middle Tertiary period, was the Dinotherium--a huge Pachyderm, twice as large as an elephant, with a tapir-like proboscis, and two great tusks curving downward from the lower jaw. It was, doubtless, aquatic in its habits, and possibly (for its hinder parts are not known), it may have been allied to the Dugong and Manatee, those whale-like Pachyderms, with a broad horizontal tail, instead of posterior limbs.

Other great herbivorous beasts roamed over the new-made land. The Mastodons, closely allied to the Elephant, had their head-quarters in North America, but extended also to Europe. And the Elephants themselves, of several species, were spread over the northern hemisphere, even to the polar regions. The Hippopotamus, the Rhinoceros, and other creatures, now exclusively tropical, were also inhabitants of the same northern latitudes.

From some specimens of Elephants and Rhinoceroses of this period, which seem to have been buried in avalanches, and thus to have been preserved from decomposition, even of the more transitory parts, as muscle and skin, we learn something of the climate that prevailed. The very fact of their preservation, by the antiseptic power of frost, shows that it was not a tropical climate in which they lived; and the clothing of thick wool, fur, and hair, which protected the skin of the Mammoth, or Siberian Elephant, tends to the same conclusion. At the same time, those regions were not so intensely cold as they are now. For the district in which the remains of Elephants and their associates are found, in almost incredible abundance, is that inhospitable coast of northern Asia which bounds the Polar Sea.

The trees of a temperate climate--the oak, the beech, the maple, the poplar, and the birch--which now attain their highest limit somewhere about 70 deg. of north latitude, and there are dwarfed to minute shrubs, appear then to have grown at the very verge of the polar basin; and that in the condition of vast and luxuriant forests, perhaps occupying sheltered valleys between mountains whose steep sides were covered with snow, already become perennial, and ever and anon rolling down in overwhelming avalanches, such as those which now occasionally descend into the valleys of the Swiss Alps.

The coast of Suffolk displays a formation known as the Crag--a local name for gravel--which rests partly on the chalk; but, as it lies in other parts over the London Clay, it is assigned to the later Tertiary, or what is called the Pleiocene period. It is divided into the coralline and the red crag, the latter being uppermost where they exist together, and therefore being the more recent. The Coralline Crag is nearly composed of corals and shells, the former almost wholly extinct now; but the latter containing upwards of seventy species still existing in the adjacent seas. The Red Crag contains few zoophytes, but is remarkable for the remains of at least five species of Whales. Other Mammalia occur in this formation, among which are the red deer and the wild boar of modern Europe.

The gradual but rapid approximation of the Tertiary fauna to that of the present surface is well indicated by Mr. Lyell's table (1841) of recent and fossil species in the English formations:--