Part 5

C D

E F

Ventral aspect of right feet of Smilisca: (A) S. baudini (KU 87177); (B) S. phaeota (KU 64276); (C) S. cyanosticta (KU 87199); (D) S. sordida (KU 91761); (E) S. puma (KU 91716), and (F) S. sila (KU 77408). x3.]

Living Smilisca: (A) S. baudini (UMMZ 115179) from 1.7 km. W Xicotencatl, Tamaulipas, Mexico; (B) S. cyanosticta (UMMZ 118163) from Volcan San Martin, Veracruz, Mexico; (C) S. phaeota (KU 64282) from Barranca del Rio Sarapiqui, Heredia Prov., Costa Rica. All approx. nat. size.]

Living Smilisca: (A) S. puma (KU 65307) from 5.9 km. W. Puerto Viejo, Heredia Prov., Costa Rica; (B) S. sila (KU 77407) from Finca Palosanto, 6 km. WNW El Volcan, Chiriqui, Panama; (C) S. sordida (KU 64257) from 20 km. WSW San Isidro el General, San Jose Prov., Costa Rica. All approx. nat. size.]

FIG. 1. Breeding site of Smilisca baudini, 4 km. WNW of Esparta, Puntarenas Prov., Costa Rica.

FIG. 2. Breeding site of Smilisca phaeota, Puerto Viejo, Heredia Prov., Costa Rica.]

FIG. 1. Breeding site of Smilisca puma, 7.5 km. W of Puerto Viejo, Heredia Prov., Costa Rica.

FIG. 2. Breeding site of Smilisca sordida, Rio La Vieja, 30 km. E of Palmar Norte, Puntarenas Prov., Costa Rica.]

Audiospectrographs and sections of breeding calls of Smilisca: (A) S. baudini (KU Tape No. 74); (B) S. cyanosticta (KU Tape No. 373); (C) S. phaeota (KU Tape No. 79).]

Audiospectrographs and sections of breeding calls of Smilisca: (A) S. puma (KU Tape No. 382); (B) S. sila (KU Tape No. 385); (C) S. sordida (KU Tape No. 398).]

Lateral views of the heads of Smilisca: (A) S. baudini (KU 87177); (B) S. sordida (KU 91765); (C) S. phaeota (KU 64276); (D) S. puma (KU 91716); (E) S. cyanosticta (KU 87199); (F) S. sila (KU 77408). x3.2.]

Aside from the skeleton referred to as Smilisca daulinia by Cope (1865:194), we have examined each of the types of the species synonymized with S. baudini. All unquestionably are representatives of S. baudini.

Taylor (1942:306) named Hyla beltrani from Tapachula, Chiapas. This specimen (UIMNH 25046) is a small female (snout-vent length, 44 mm.) of S. baudini. Taylor (1954:630) named Hyla manisorum from Batan, Limon, Costa Rica. The type (KU 34927) is a large female (snout-vent length, 75.3 mm.) S. baudini. In this specimen and a male from Suretka, Costa Rica, the usual dorsal color pattern is absent, but the distinctive curved supraorbital processes, together with other structural features, show that the two specimens are S. baudini.

Hyla baudini dolomedes Barbour (1923:11), as shown by Dunn (1931a:413), was based on a specimen of Smilisca phaeota from Rio Esnape, Darien, Panama.

Distribution.--Smilisca baudini inhabits lowlands and foothills usually covered by xerophytic vegetation or savannas, but in the southern part of its range baudini inhabits tropical evergreen forest. The species ranges throughout the Pacific and Atlantic lowlands of Mexico from southern Sonora and the Rio Grande embayment of Texas southward to Costa Rica, where on the Pacific lowlands the range terminates at the southern limits of the arid tropical forest in the vicinity of Esparta; on the Caribbean lowlands the distribution seems to be discontinuous southward to Suretka (Fig. 1). Most localities where the species has been collected are at elevations of less than 1000 meters. Three localities are notably higher; calling males were found at small temporary ponds in pine-oak forest at Linda Vista, 2 kilometers northwest of Pueblo Nuevo Solistahuacan, Chiapas, elevation 1675 meters, and 10 kilometers northwest of Comitan, Chiapas, at an elevation of 1925 meters. Tadpoles and metamorphosing young were obtained from a pond in arid scrub forest, 17 kilometers northeast of Juchatengo, Oaxaca, elevation 1600 meters. Stuart (1954:46) recorded the species at elevations up to 1400 meters in the south-eastern highlands of Guatemala.

Specimens examined.--3006, as follows: UNITED STATES: TEXAS: Cameron County, Brownsville, CNHM 5412-3, 6869, UMMZ 54036, USNM 3256.

MEXICO: =Campeche=: Balchacaj, CNHM 102285, 102288, 102291, 102311, UIMNH 30709-22, 30726; Champoton, UMMZ 73172 (2), 73176, 73180; 16 km. E Champoton, UMMZ 73181; 5 km. S Champoton, KU 71369-75; 9 km. S Champoton, KU 71367-8; 10.5 km. S Champoton, KU 71365-6, 71722 (tadpoles), 71723 (yg.); 24 km. S Champoton, UMMZ 73177 (2); Chuina, KU 75101-3; Ciudad del Carmen, UIMNH 30703-8; Dzibalchen, KU 75413-31; Encarnacion, CNHM 102282, 102289, 102294-5, 102300, 102306-8, 102312, 102314, 102316-7, 102319, 102322, UIMNH 30727-40, 30836-7; 1 km. W Escarcega, KU 71391-6; 6 km. W Escarcega, KU 71397-403; 7.5 km. W Escarcega, KU 71376-89; 14 km. W Escarcega, KU 71390; 13 km. W, 1 km. N Escarcega, KU 71404; 3 km. N Hopelchen, KU 75410-11; 2 km. NE Hopelchen, KU 75412; Matamoras, CNHM 36573; Pital, UIMNH 30741; 1 km. SW Puerto Real, Isla del Carmen, KU 71345-64; San Jose Carpizo, UMMZ 99879; Tres Brazos, CNHM 102284, UIMNH 30723-5; Tuxpena Camp, UMMZ 73239.

=Chiapas=: Acacoyagua, USNM 114487-92; 2 km. W Acacoyagua, USNM 114493-4; 5 km. E Arroyo Minas, UIMNH 9533-7; Berriozabal, UMMZ 119186 (7); Chiapa de Corzo, UMMZ 119185 (2); Cintalapa, UIMNH 50077; Colonia Soconusco, USNM 114495-9; 5 km. W Colonia Soconusco, UMMZ 87885 (7); Comitan, UMMZ 94438; 10 km. NW Comitan, KU 57185; El Suspiro, UMMZ 118819 (11); Escuintla, UMMZ 88271 (7), 88278, 88327, 109233; 6 km. NE Escuintla, UMMZ 87856 (26); 3 km. E Finca Juarez, UIMNH 9538; Finca Prussia, UMMZ 95167; Honduras, UMMZ 94434-7; La Grada, UMMZ 87862; 21 km. S La Trinitaria, UIMNH 9540-1; 14.4 km. SW Las Cruces, KU 64239-44; Palenque, UIMNH 49286, USNM 114473-84; 2 km. NW Pueblo Nuevo Solistahuacan, KU 57182-4, UMMZ 119948 (8), 121514; 1.3 km. N Puerto Madero, KU 57186-9; 4 km. N Puerto Madero, KU 57190-1; 8 km. N Puerto Madero, UMMZ 118379 (2); 12 km. N Puerto Madero, KU 57192; 17.6 km. N Puerto Madero, UMMZ 118378; Rancho Monserrata, UIMNH 9531-2, UMMZ 102266-7; Region Soconusco, UIMNH 33542-56; San Bartola, UIMNH 9519-30; San Geronimo, UIMNH 30804; San Juanito, USNM 114485-6; San Ricardo, CNHM 102406; Solosuchiapa, KU 75432-3; Tapachula, CNHM 102208, 102219, 102239, 102405, UIMNH 25046, 30802-3; Tonola, AMNH 531, CNHM 102232, 102416, UIMNH 30805-9, USNM 46760; Tuina, KU 41593 (skeleton); Tuxtla Gutierrez, CNHM 102231, 102248; 6 km. E Tuxtla Gutierrez, UIMNH 9539; 10 km. E Tuxtla Gutierrez, UMMZ 119949.

=Chihuahua=: 2.4 km. SW Toquina, KU 47226-7; Riito, KU 47228.

=Coahuila=: mountain near Saltillo, UIMNH 30833-4.

=Colima=: No specific locality, CNHM 1632; Colima, AMNH 510-11; Hacienda Albarradito, UMMZ 80029 (2); Hacienda del Colomo, AMNH 6208; Los Mezcales, UMMZ 80028; Manzanillo, AMNH 6207, 6209; Paso del Rio, CNHM 102207, 102229-30, UIMNH 30819-21, UMMZ 110875 (3); Periquillo, UMMZ 80025 (3), 80026 (14); 1.6 km. SW Pueblo Juarez, UMMZ 115564; Queseria, CNHM 102204, 102216-7, 102224, UIMNH 30816-8, UMMZ 80023 (7), 80024 (7); Santiago, UMMZ 80027; 7.2 km. SW Tecolapa, UMMZ 115184.

=Guerrero=: Acahuizotla, UF 1338 (2), 1339-40, UMMZ 119182 (2), 119184; 3 km. S Acahuizotla, KU 87183-7; Acapulco, AMNH 55276, UMMZ 121879 (4), USNM 47909; 3 km. N Acapulco, UMMZ 110127; 8 km. NW Acapulco, UF 11203 (7); 27 km. NE Acapulco, UIMNH 26597-610; Agua del Obispo, CNHM 102214, 102290, 102293, 102310, 102413, KU 60413, 87180-2, UIMNH 30764-6; Atoyca, KU 87175-8; Buena Vista, CNHM 102279, 102304, 102313, 102315, UIMNH 30774; Caculutla, KU 87179; 20 km. S Chilpancingo, CNHM 102242, 102401, 102410-1, 102415; Colonia Buenas Aires, UMMZ 119189; El Limoncito, CNHM 102292, 102303, 102321, 102414; El Treinte, CNHM 102212, 102221, 102237, 102240-1, UIMNH 30783-5, USNM 114508-10; Laguna Coyuca, UMMZ 80960 (2); 3 km. N Mazatlan, UIMNH 30777-9; 9 km. S Mazatlan, CNHM 102209, 102215, 102234, 102246, UIMNH 30781-2; Mexcala, CNHM 102399, 102403, 102409, 106539-40, UIMNH 30775-6; Ocotito, KU 60414-23; 5.4 km. N Ocotito, UMMZ 119181 (4); 1.6 km. N Organos, UIMNH 30752-63; Palo Blanco, CNHM 102283, 102286, 102305, 102320, 102404, UIMNH 30767-70; Pie de la Cuesta, AMNH 55275, 59202-5; Puerto Marquez, AMNH 59200-1 (13); 5.6 km. S San Andreas de la Cruz, KU 87173-4; San Vincente, KU 87172; Zaculapan, UMMZ 119183.

=Hidalgo=: Below Tianguistengo, CNHM 102318.

=Jalisco=: Atenqueque, KU 91435-6; 5 km. NE Autlan, UIMNH 30810; 5 km. E Barro de Navidad, UMMZ 110900; Charco Hondo, UMMZ 95247; Puerto Vallarta, UIMNH 41346; between La Huerta and Tecomates, KU 91437; 3 km. SE La Resolana, KU 27619, 27620 (skeleton); 11 km. S, 1.6 km. E Yahualica, KU 29039; Zapotilitic, CNHM 102238.

=Michoacan=: Aguililla, UMMZ 119179 (5); Apatzingan, CNHM 38766-90, KU 69101 (skeleton); 7 km. E Apatzingan, UMMZ 112843; 11 km. E Apatzingan, UMMZ 112841 (3); 27 km. S Apatzingan, KU 37621-3; 1.6 km. N Arteaga, UMMZ 119180; Charapendo, UMMZ 112840; Coahuayana, UMMZ 104458; El Sabino, CNHM 102205-6, 102210-1, 102220, 102228, 102233, UIMNH 30822-3; La Placita, UMMZ 104456; La Playa, UMMZ 105163; 30 km. E Nueva Italia, UMMZ 120255 (2); 4 km. S Nueva Italia, UMMZ 112842; Ostula, UMMZ 104457 (4); Salitre de Estopilas, UMMZ 104459; San Jose de la Montana, UMMZ 104461 (2); 11 km. S Tumbiscatio, KU 37626; 12 km. S Tzitzio, UMMZ 119178.

=Morelos=: 3.5 km. W Cuautlixco, KU 87188-90; 1 km. NE Puente de Ixtla, KU 60393-4; 20 km. S Puente de Ixtla, CNHM 102400, UIMNH 30832; Tequesquitengo, AMNH 52036-9.

=Nayarit=: 3 km. S Acaponeta, UMMZ 123030 (4); 56 km. S Esquinapa (Sinaloa), KU 73909; Jesus Maria, AMNH 58239; San Blas, KU 28087, 37624, 62360-2, USNM 51408; 8.6 km. E San Blas, UMMZ 115185; Tepic, UIMNH 30812-5; 4 km. E Tuxpan, KU 67786; 11 km. SE Tuxpan, UIMNH 7329-31, 7335-59.