Part 3

Snout long, sloping, rounded; vocal sacs in breeding males white; cream-colored or pale blue flecks on flanks and posterior surfaces of thighs S. sordida, p. 323

Key to Tadpoles

1. Pond tadpoles; tail about half again as long as body; mouth anteroventral 2

Stream tadpoles; tail about twice as long as body; mouth ventral 5

2. Labial papillae in two rows 3

Labial papillae in one row 4

3. First upper tooth row strongly arched medially; third lower tooth row much shorter than other rows; dorsal fin deepest at about two-thirds length of tail; tail cream-colored with dense gray reticulations S. puma, p. 314

First upper tooth row not arched medially; third lower tooth row nearly as long as others; dorsal fin deepest at about one-third length of tail; tail tan with brown flecks and blotches S. baudini, p. 289

4. Dorsal fin extending onto body S. phaeota, p. 308

Dorsal fin not extending onto body S. cyanosticta, p. 303

5. Mouth completely bordered by two rows of papillae; inner margin of upper beak not forming continuous arch with lateral processes; red or reddish brown markings on tail S. sordida, p. 323

Median part of upper lip bare; rest of mouth bordered by one row of papillae; inner margin of upper beak forming continuous arch with lateral processes; dark brown markings on tail S. sila, p. 318

ACCOUNTS OF SPECIES

=Smilisca baudini= (Dumeril and Bibron)

Hyla baudini Dumeril and Bibron, Erpetologie general, 8:564, 1841 [Holotype.--MNHN 4798 from "Mexico;" Baudin collector]. Guenther, Catalogue Batrachia Salientia in British Museum, p. 105, 1858. Brocchi, Mission scientifique au Mexique ..., pt. 3, sec. 2, Etudes sur les batrachiens, p. 29, 1881. Boulenger, Catalogue Batrachia Salientia in British Museum, p. 371, Feb. 1, 1882. Werner, Abhand. Zool.-Bot. Gesell. Wien., 46:8, Sept. 30, 1896. Guenther, Biologia Centrali-Americana: Reptilia and Batrachia, p. 270, Sept. 1901. Werner, Abhand. Konigl. Akad. Wiss. Munchen, 22:351, 1903. Cole and Barbour, Bull. Mus. Comp. Zool., 50(5):154, Nov. 1906. Gadow, Through southern Mexico, p. 76, 1908. Ruthven, Zool. Jahr. 32(4):310, 1912. Decker, Zoologica, 2:12, Oct., 1915. Stejneger and Barbour, A checklist of North American amphibians and reptiles, p. 32, 1917. Noble, Bull. Amer. Mus. Nat. Hist., 38(10):341, June 20, 1918. Nieden, Das Tierreich, Amphibia, Anura I, p. 243, June, 1923. Gadow, Jorullo, p. 54, 1930. Dunn and Emlen, Proc. Acad. Nat. Sci. Philadelphia, 84:24, March 22, 1932. Kellogg, Bull. U. S. Natl. Mus., 160:160, March 31, 1932. Martin, Aquarien Berlin, p. 92, 1933. Stuart, Occas. Papers Mus. Zool., Univ. Michigan, 292:7, June 29, 1934; Misc. Publ. Mus. Zool. Univ. Michigan, 29:38, Oct. 1, 1935. Gaige, Carnegie Inst. Washington, 457:293, Feb. 5, 1936. Gaige, Hartweg, and Stuart, Occas. Papers Mus. Zool. Univ. Michigan, 360:5, Nov. 20, 1937. Smith, Occas. Papers Mus. Zool. Univ. Michigan, 388:2, 12, Oct. 31, 1938; Ann. Carnegie Mus., 27:312, March 14, 1939. Taylor, Copeia, 2:98, July 12, 1939. Hartweg and Oliver, Misc. Publ. Mus. Zool. Univ. Michigan, 47:12, July 13, 1940. Schmidt and Stuart, Zool. Ser. Field Mus. Nat. Hist., 24(21):238, August 30, 1941. Schmidt, Zool. Ser. Field Mus. Nat. Hist., 22(8):486, Dec. 30, 1941. Wright and Wright, Handbook of frogs and toads, Ed. 2, p. 134, 1942. Stuart, Occas. Papers Mus. Zool. Univ. Michigan, 471:15, May 17, 1943. Bogert and Oliver, Bull. Amer. Mus. Nat. Hist., 83(6):343, March 30, 1945. Taylor and Smith, Proc. U. S. Natl. Mus., 95(3185): 590, June 30, 1945. Smith, Ward's Nat. Sci. Bull., 1, p. 3, Sept., 1945. Schmidt and Shannon, Fieldiana, Zool. Chicago Nat. Hist. Mus., 31(9):67, Feb. 20, 1947. Stuart, Misc. Publ. Mus. Zool. Univ. Michigan, 69:26, June 12, 1948. Wright and Wright, Handbook of frogs and toads, Ed. 3, p. 298, 1949. Stuart, Contr. Lab. Vert. Biol. Univ. Michigan, 45:22, May, 1950. Mertens, Senckenbergiana, 33:170, June 15, 1952; Abhand. Senckenb. Naturf. Gesell., 487:28, Dec. 1, 1952. Schmidt, A checklist of North American amphibians and reptiles, Ed. 6, p. 69, 1953. Stuart Contr. Lab. Vert. Biol. Univ. Michigan, 68:46, Nov. 1954. Zweifel and Norris, Amer. Midl. Nat., 54(1):232, July 1955. Martin, Amer. Nat., 89:356, Dec. 1955. Duellman, Copeia, 1:49, Feb. 21, 1958. Goin, Herpetologica, 14:119, July 23, 1958. Turner, Herpetologica, 14:192, Dec. 1, 1958. Conant, A field guide to reptiles and amphibians, p. 284, 1958. Duellman, Univ. Kansas Publ., Mus. Nat. Hist., 13(2):59, Aug. 16, 1960; Univ. Kansas Publ., Mus. Nat. Hist., 15(1): 46, Dec. 20, 1961. Porter, Herpetologica, 18:165, Oct. 17, 1962.

Hyla vanvlietii Baird, Proc. Acad. Nat. Sci. Philadelphia, 7:61, April 27, 1854 [Holotype.--USNM 3256 from Brownsville, Cameron County, Texas; S. Van Vliet collector]. Baird, United States and Mexican boundary survey, 2:29, 1859. Smith and Taylor, Univ. Kansas Sci. Bull., 33:361, March 20, 1950. Cochran, Bull. U. S. Natl. Mus., 220:60, 1961.

Hyla vociferans Baird, United States and Mexican boundary survey, 2:35 1859 [nomen nudum]. Diaz de Leon, Indice de los batracios que se encuentran en la Republica Mexicana, p. 20, June 1904.

Hyla muricolor Cope, Proc. Acad. Nat. Sci. Philadelphia, 14(9):359, 1862 [Holotype.--USNM 25097 from Mirador, Veracruz, Mexico; Charles Sartorius collector]. Smith and Taylor, Univ. Kansas Sci. Bull., 33:349, March 20, 1950. Cochran, Bull. U. S. Natl. Mus., 220:56, 1961.

Smilisca daulinia Cope, Proc. Acad. Nat. Sci. Philadelphia, 17:194, Oct. 1865 [Holotype.--"skeleton in private anatomical museum of Hyrtl, Professor of Anatomy in the University of Vienna"]. Smith and Taylor, Univ. Kansas Sci. Bull., 33:347, March 20, 1950.

Smilisca daudinii [lapsus for baudini], Cope, Proc. Acad. Nat. Sci. Philadelphia, 23, pt. 2:205, 1871.

Smilisca baudini, Cope, Bull. U. S. Nat. Mus., 1:31, 1875; Jour. Acad. Nat. Sci. Philadelphia, 8, pt. 2:107, 1876; Proc. Amer. Philos. Soc., 18:267, August 11, 1879. Yarrow, Bull. U. S. Nat. Mus., 24:176, July 1, 1882. Cope, Bull. U. S. Nat. Mus., 32:13, 1887; Bull. U. S. Nat. Mus., 34:379, April 9, 1889. Dickerson, The frog book, p. 151, July, 1906. Smith and Taylor, Univ. Kansas Sci. Bull., 33:442, March 20, 1950; Taylor, U. Kan. Sc. Bull., 34:802, Feb. 15, 1952; Univ. Kansas Sci. Bull., 35:794, July 1, 1952. Brattstrom, Herpetologica, 8(3):59, Nov. 1, 1952. Taylor, U. Kan. Sci. Bull., 35:1592, Sept. 10, 1953. Peters, Occas. Papers Mus. Zool. Univ. Michigan, 554:7, June 23, 1954. Duellman, Occas. Papers Mus. Zool. Univ. Michigan, 560:8, Oct. 22, 1954. Chrapliwy and Fugler, Herpetologica, 11:122, July 15, 1955. Smith and Van Gelder, Herpetologica, 11:145, July 15, 1955. Lewis and Johnson, Herpetologica, 11:178, Nov. 30, 1955. Martin, Misc. Publ. Mus. Zool. Univ. Michigan, 101:53, April 15, 1958. Stuart, Contr. Lab. Vert. Biol. Univ. Michigan, 75:17, June, 1958. Minton and Smith, Herpetologica, 17:74, July 11, 1961. Nelson and Hoyt, Herpetologica, 17:216, Oct. 9, 1961. Holman, Copeia, 2:256, July 20, 1962. Stuart, Misc. Publ. Mus. Zool. Univ. Michigan, 122:41, April 2, 1963. Maslin, Herpetologica, 19:124, July 3, 1963. Holman and Birkenholz, Herpetologica, 19:144, July 3, 1963. Duellman, Univ. Kansas Publ. Mus. Nat. Hist., 15(5):228, Oct. 4, 1963. Zweifel, Copeia, 1:206, March 26, 1964. Duellman and Klaas, Copeia, 2:313, June 30, 1964. Davis and Dixon, Herpetologica, 20:225, January 25, 1965. Neill, Bull. Florida State Mus., 9:89, April 9, 1965.

Hyla pansosana Brocchi, Bull. Soc. Philom., ser. 7, 1:125, 1877 [Holotype.--MNHN 6313 from Panzos, Alta Verapaz, Guatemala; M. Bocourt collector]; Mission scientifique au Mexique ..., pt. 3, sec. 2, Etudes sur les batrachiens, p. 34, 1881.

Hyla baudini baudini, Stejneger and Barbour, A checklist of North American amphibians and reptiles, Ed. 3, p. 34, 1933. Wright and Wright, Handbook of frogs and toads, p. 110, 1933. Stejneger and Barbour, A checklist of North American amphibians and reptiles, Ed. 4, p. 39, 1939; A checklist of North American amphibians and reptiles, Ed. 5, p. 49, 1943. Smith and Laufe, Trans. Kansas Acad. Sci., 48(3):328, Dec. 19, 1945. Peters, Nat. Hist. Misc., 143:7, March 28, 1955.

Hyla beltrani Taylor, Univ. Kansas Sci. Bull. 28(14):306, Nov. 15, 1942 [Holotype.--UIMNH 25046 (formerly EHT-HMS 29563) from Tapachula, Chiapas, Mexico; A. Magana collector]. Smith and Taylor, Bull. U. S. Natl. Mus. 194:87, June 17, 1948; Univ. Kansas Sci. Bull, 33:326, March 20, 1950. Smith, Illinois Biol. Mono., 32:23, May, 1964.

Smilisca baudini baudini, Smith, Jour. Washington Acad. Sci., 37(11):408, Nov. 15, 1947. Smith and Taylor, Bull. U. S. Natl. Mus., 194:75, June 17, 1948; Univ. Kansas Sci. Bull., 33:347, March 20, 1950. Brown, Baylor Univ. Studies, p. 68, 1950. Smith, Smith, and Werler, Texas Jour. Sci., 4(2):254, June 30, 1952. Smith and Smith, Anales Inst. Biol., 22(2):561, Aug. 7, 1952. Smith and Darling, Herpetologica, 8(3):82, Nov. 1, 1952. Davis and Smith, Herpetologica, 8(4):148, Jan. 30, 1953. Neill and Allen, Publ. Res. Div. Ross Allen's Reptile Inst., 2(1):26, Nov. 10, 1959. Maslin, Univ. Colorado Studies, Biol. Series, 9:4, Feb. 1963. Holman, Herpetologica, 20:48, April 17, 1964.

Hyla manisorum Taylor, Univ. Kansas Sci. Bull., 36:630, June 1, 1954 [Holotype.--KU 34927 from Batan, Limon Province, Costa Rica; Edward H. Taylor collector]. Duellman and Berg, Univ. Kansas Publ. Mus. Nat. Hist, 15(4):193, Oct. 26, 1962.

Diagnosis.--Size large ( 76 mm., 90 mm.); skull noticeably wider than long, having small frontoparietal fontanelle (roofed with bone in large individuals); postorbital processes long, pointed, curving along posterior border of orbit; squamosal large, contacting maxillary; tarsal fold strong, full length of tarsus; inner metatarsal tubercle large, high, elliptical; hind limbs relatively short, tibia length less than 55 per cent snout-vent length; lips strongly barred with brown and creamy tan; flanks pale cream with bold brown or black reticulations in groin; posterior surfaces of thighs brown with cream-colored flecks; dorsal surfaces of limbs marked with dark brown transverse bands. (Foregoing combination of characters distinguishing S. baudini from any other species in genus.)

Description and Variation.--Considerable variation in size, and in certain proportions and structural characters was observed; variation in some characters seems to show geographic trends, whereas variation in other characters apparently is random. Noticeable variation is evident in coloration, but this will be discussed later.