If we confine ourselves, however, to the conclusions regarding ancient man which are generally accepted, we must say that he has not been clearly traced back beyond the Glacial Period, though some of the relics found in the older river gravels and in the lowest cave accumulations may well be of pre-glacial age. Many geologists believe that he reached Europe as early as the extinct mammals with which he was contemporaneous there, but how far back in time this would carry his advent it is impossible to say.
Coming now to the consideration of more immediate human relics, the bones of man himself, it must be said that well-authenticated remains of palæolithic or early neolithic man are not numerous. As long as man left his bones to the unaided agencies of nature, they were little likely to be preserved. Of the anthropoid apes of Europe, probably numerous in individuals, a few remains of one or two species alone survive. Of pre-glacial man none remain, but this may merely indicate that he has shared the fate of numerous other species that died out and left no trace. It was only when the growing cold drove man from the open woods to seek shelter in caves that remnants of his body were likely to be preserved, and only when a growing sense of human dignity led to the art of sepulture that the preservation of his bones became assured.
The burial art was seemingly not practised by the hunters of the river-drift period or by men of still earlier date. The only remains of primitive man known are those found in caves and rock shelters. A number of human skulls have been discovered in these situations, and in a few instances skeletons have been exhumed. In the neolithic period interment became more common and more carefully performed, and the progress of this period is marked by many remains of man, which in later times were buried in elaborately constructed stone sepulchres, sometimes massive in materials and covered by great earth-mounds.
What is meant by the Glacial Age is probably well-known to most readers, but its close relations to ancient man render it important for those who are not familiar with its meaning that a passing description of it should here be given. It will suffice to say that there are found over much of the northern portions of America and Europe accumulations of clays, sands, and gravels, sometimes laid down in stratified beds, sometimes rudely piled together. In these occur blocks of stone, large and small, and other blocks, occasionally of great size, are found in isolated localities. The solid rocks which lie beneath these heaps are often scratched or polished, as if the material had been pushed over them with great force.
All geologists now believe that these accumulations were made by ice, at some remote period when a very cold climate prevailed in the northern hemisphere, and great glaciers slowly made their way southward, grinding and rending as they went, and burying the land under their mountain-like heaps, which sometimes were a mile or more in depth. In North America the glacial ice pushed southward to the 40th degree of north latitude. In Europe it extended to the Alpine region, but failed to reach the countries bordering on the Mediterranean.
The elaborate and minute investigation of the glacial deposits has made it highly probable that there were two glacial eras, two periods in which the ice pushed down far to the south, and that these were separated by a period in which the ice retreated and an age of warmer weather intervened. This is known as the interglacial period. So far as can be positively ascertained, all the authentic relics of man belong to the Glacial Age. They seem first to become numerous in the interglacial period, and continue to increase and become diversified as we descend lower in time. How long ago it was that the sea of ice began its downflow over the earth it is impossible to say. Some place it back six hundred thousand or seven hundred thousand years. Some seek to bring it down to a quite recent date. It is still so uncertain and such a matter of controversy that the utmost we are able definitely to say is that it was very long ago.
While there is no positive proof that men dwelt in Europe before the coming on of the glacial chill, we have no just reason to doubt it. That he lived there during glacial times is unquestionable, and we may be very well assured that a naked tropical animal, destitute of the hairy covering of the other animals, would not have chosen that frozen period to migrate to the north. The fact that he was there during the ice age seems satisfactory evidence that he was there before that age, during the mild climate of late Tertiary times, and that--for a reason which we shall hereafter consider--he was caught there and unable to retreat, and was forced to adapt himself to the new conditions.
During the warm preceding period he probably wandered as a hunter through the European forests. But with the gradual coming on of a wintry chill, as the advance of the ice began, shelter of some kind became necessary, and he sought refuge in caves. From being a forest wanderer he became a troglodyte. Everywhere in southwestern Europe we find traces of this period of man's existence. There is hardly a cave or rock shelter in that region within which he has not left his marks. He made his way to England, which was probably then connected by land with Europe, and dwelt long in its caverns. His period of cave residence, indeed, appears to have been a very extended one. While it continued, deposits many feet in depth gradually accumulated on the floors of the caverns, slowly filling them up. And that, in some cases at least, this cave residence ended a very long time ago, we are assured, for since then a great thickness of stalagmite, which is deposited with extreme slowness, has spread over the lower cave deposits and sealed them in.
It is in these caves that we find, not only the rude stone spearheads, scrapers, hammers, etc., the bone awls, borers, and other implements of palæolithic man, but the bones of man himself. And it is significant of his primitive condition that these earliest relics indicate a man of a very low grade of development, mentally far above the ape, it is true, but mentally and physically much below modern man.
The most ape-like of those human remains is the famous Neanderthal skull, found in 1856 in a limestone cavern of the Neanderthal Valley, between Düsseldorf and Elberfeld, in Rhenish Prussia. The relics discovered consist of the brain cap, two femori, two humeri, and other fragments. The fragment of the skull attracted wide attention by its bestial aspect, it presenting a low, narrow and receding forehead, and an enormous thickness of the bony ridges over the eyes, like that seen in the gorilla. This skull, which was associated with remains of the cave-bear, hyena, and rhinoceros, is, with one exception, the most ape-like human relic yet found. Yet its cranial capacity is far above that of the highest apes, and is assimilated with that of Hottentot and Polynesian skulls.
It has been maintained that this is a pathological specimen, and does not represent normal man. But this theory has been disproved by the fact that other skulls of similar cranial characters are now known, indicating that the Neanderthal cranium represents a type of man, not an abnormal individual. In the Spy Cavern, in the province of Namur, Belgium, there were found, in 1886, two nearly perfect skeletons of a man and a woman, both of them with very prominent eye ridges, low, retreating foreheads, and large orbits. This was strikingly the case with the woman. The lower jaws in both were heavy, while the woman was almost destitute of a chin--a marked ape-like characteristic. The tibia was shorter than in any known race and stouter than in most. Its curious feature was the articulation with the femur, which was such that to maintain the equilibrium the head and body must have been thrown forward, as is the case in the anthropoid apes.
In the cave of Naulette, near Dinant, Belgium, has been found the lower jaw of a man of decidedly ape-like aspect. Its prognathism or protrusion is extreme, and the canine teeth were very strong, while the molars were evidently large and increased in size backward, a non-human characteristic. At La Denise, in the upper Loire, France, have been found the frontal bones of a man like the Neanderthal man in type, the forehead being depressed and retreating, and the superciliary ridges large and thick. Several other skulls of this general type are known, but the above will suffice as examples.
Remains of palæolithic man of considerably higher type are not wanting. In the rock shelter of Cro-Magnon, France, were found the bones of three men, one woman, and one child, of more advanced character. These, however, are of late date and may have been early neolithic. At Engis, near Liège, Belgium, a deeply buried skull, associated with many remains of extinct animals, has been dug up, which is by no means ape-like in character. A still superior example of palæolithic man is the skeleton found in a cavern at Mentone, east of Nice, France, which represents a man six feet in height, with rather large head, high forehead, and very large facial angle (85°). The cave contained bones of extinct animals, but no trace of the reindeer.
There is no occasion to speak here of the many remains of neolithic man that have been exhumed. Sparse in the early part of the age of polished stone weapons, they gradually became numerous, and merged into the human remains of late prehistoric times. The American continent is not without its relics of ancient man, the most famous of which is the Calaveras skull, found in 1886 in the auriferous gravels of Calaveras County, California, at an extraordinary depth. The miners, in excavating a shaft, passed through several layers of lava and gravel, forming a total thickness of seventy-nine feet of lava and a considerable thickness of gravel, making nearly one hundred and thirty feet in all. At this depth a skull was found imbedded in the gravel, which, if authentic, must have been overflowed by several successive thick outpours of lava in the ancient volcanic era of that region. As its authenticity is, however, still a matter of controversy, nothing further need here be said about it.
Leaving these evidences of human antiquity, we come to the most remarkable and significant of all the known relics of man, if indeed it is man, for it seems to many a link between man and the ape,--not yet human, while no longer simian. This is the fossil find made by Dr. Eugene Dubois in 1891 on the banks of the Bengawan River, Java, and named by him Pithecanthropus erectus, he maintaining that it represents a new genus of upright animals, or even a new family. The remains found by him consisted of the upper part of a skull, a molar tooth, and a femur, possibly not belonging to a single individual, as they were somewhat separated. These were exhumed from a stratum of volcanic tufa, claimed to be of Tertiary age, but perhaps Quaternary, and lay at a depth of some forty feet beneath the surface.
The femur very closely resembles that of a human being of average size, and its shape, articulating surface, and other characters show clearly that the animal stood habitually erect. The principal significance lies in the tooth and the cranium. The former is like that of the chimpanzee in shape, but less rugose on its grinding surface. It seems to lie between the ape and the human type of dentition. The cranium has a low, depressed arch, with a very narrow frontal region and highly developed superciliary ridges. The cranial capacity was apparently about one thousand, that of man being from thirteen hundred to fourteen hundred. It is therefore said to be "the lowest human cranium yet described, very nearly as much below the Neanderthal as that is below the normal European."
Professor O. C. Marsh, in a paper on the subject in the American Journal of Science, for February, 1895, agrees with Dr. Dubois in his view of the distinct position of this form in the animal kingdom, and says that the discoverer "has proved the existence of a new prehistoric anthropoid form, not human, indeed, but in size, brain power, and erect posture much nearer man than any animal hitherto discovered, living or extinct."
We have here given a short review of a long story. The evidences of man's former existence upon the earth are multitudinous, but any extended consideration of them is aside from our purpose, which is merely to show that the proofs of man's descent found in his physical structure are strengthened by evidences which he has left strewn behind him in his long march down the ages. Only a single conclusion can be drawn from these vestiges of man excavated from caves and gravels, namely, that they indicate a gradual and steady progression upward from a very low condition, while they nowhere give evidence of the traditional fall of man.
This is certainly the case with the relics of human workmanship. They begin with the rudest chipped stones, and very slowly improve in form and finish and become more varied, as we move upward in our search. The ground and polished stones follow, and the variety of implements considerably increases, until at length the age of metal, with its developed industries, is reached. The only seeming evidence of superior intellect to be found in this gradual progress is that of the drawings and carvings left us by one group of palæolithic men. But the actual mental development indicated by these becomes problematical when we consider that similar drawings are made to-day by the Bushmen of South Africa, a race of men occupying a very low mental stage. From this fact we may fairly conclude that the possession of a simple graphic art does not necessarily indicate any considerable intellectual advance.
If we consider the remains of man himself, the few bones which mark his early pathway through time, a similar conclusion must be drawn. Beginning with Pithecanthropus, which science is yet in doubt whether to class with the apes or with men, we pass upward to the bestial Neanderthal man and his fellows of the same low type. Of the sparse remains of palæolithic man that exist, the most are of this degraded type. The cranial capacity is usually not small. They had the full brain development of man. But this simply assimilates them with the low races of existing savages, many of whom have not developed the simple art of chipping stone to form weapons and yet have brains of normal human weight.
In truth, the influences under which the development of the brain took place were not what we now call intellectual. Developing man used his mental powers actively in his dealings with the hostile forces of surrounding nature, and nearly all the forces of evolution were brought to bear upon the organ of the mind, the body remaining practically unchanged. His senses became acute, his cunning and alertness high, his use of weapons skilful, but his field of mental exercise was still the outer world, and the inner world of thought remained in its embryo state. The more recent development of the mind has been in its intellectual powers, while its physical aptitudes have somewhat declined. This has not yielded any marked increase in the dimensions of the brain, but it may have had a decided effect upon the proportion of its parts, the regions of the cerebrum devoted to intellectual activity probably increasing at the expense of the motor and sensory regions, while the convolutions may have grown considerably more complicated.
FROM QUADRUPED TO BIPED
In the question which now confronts us, that of the evolution of man from the lower world of animals, it is necessary first to state in what particulars he has evolved, what are the conditions which distinguish him from the lower animals. Four marked distinctions may be named: his erect attitude, with the freeing of the fore limbs from use as agents in locomotion; his employment of natural objects, instead of his bodily organs, as tools and weapons; his development of vocal language; and his great mental superiority, with the general use of the mind in his dealings with nature.
In none of these particulars does man stand quite alone; in all of them an affinity with the lower animals exists. Steps of progress in these directions have been made by many animals, though none of them have gained any considerable advance. In man's strikingly developed social habit and organization he has no close counterpart among the vertebrates, but several among the insects. And it is of much interest to find that in the highest field of man's progress, his employment of the mind in his dealings with nature, he is chiefly emulated by such lowly-organized creatures as the ants and the bees.
We do not need to look far among the lower animals for the species which come nearest to man in structure and which seem to have immediately preceded him in the line of descent. We find these forms in the monkeys or apes, and especially in their highest representatives, the anthropoid apes. These possess in a partial degree all the special characteristics of man. They are social in habit; some of them are semi-erect in posture, and their fore limbs partly freed from use in locomotion; they possess some imperfect means of vocal communication; they employ the mind to some extent in place of the body; in short, they seem arrested forms on the road from brute to man, signal-posts on the highway of evolution. In physical organization their approach to man is singularly close. In anatomy man and the higher apes are in most respects counterparts of each other. The principal anatomical distinction has been considered to be in the foot, which from the opposable character of the great toe was classed by Cuvier with the hand, the apes being named Quadrumana, or four-handed, and man Bimana, or two-handed. Fuller research has shown that this distinction does not exist, the foot of the ape being found to agree far more closely with the foot than with the hand of man. Estimated according to use, the hand is, in the whole order, the special prehensile organ; the foot, however prehensile it may be, is predominantly a walking organ. And the opposability of the great toe is approached in some men, who have great mobility in this organ, and can use it for grasping.
In regard to the brain, the organ of the mind, the difference between the higher apes and man is almost solely one of comparative size, the lower intelligence of the apes being indicated by the smaller size of their brains. The largest ape brain is scarcely half the size of the smallest human brain. But anatomically they are nearly identical. All the structural features of the brain are common to both, and the details are largely filled out in the anthropoid apes, the convolutions being all present and the pattern of arrangement the same. The brain of the orang may be said to be like that of man in all respects except size and the greater symmetry of its convolutions, which are less complicated with minor convolutions than in man. In truth, the difference between the brains of man and the orang is almost insignificant as compared with the difference between those of the orang and the lowest apes. Mr. E. W. Taylor, who has recently made an exhaustive study of the minute anatomy of the brain of the chimpanzee, remarks, "The similarity between the brain of the anthropoid apes and of man is one of the most singular and interesting facts of which we have knowledge."
In any attempt, then, to consider the origin of man from the point of view of evolution, we are irresistibly drawn to the ape tribe as the next lower link in the long chain of development, and are led to consider the characteristics of the apes as the intermediate stage between the quadruped and the biped, the bridge crossing this great gulf in organic development. This is by no means to suggest that some one of the existing anthropoid apes is the direct ancestor of man. Such an idea has never been entertained by scientists. These animals cannot even fairly be considered as brothers to man's ancestor, but must be looked upon as more or less distant cousins, with a physical organization less favorable to high development than that of man. Man's ancestry lies much farther back in time, and his progenitor must have been constituted differently from any of the existing large apes.
In the ape tribe we are able to trace nearly every step by which the gulf between quadruped and biped has been crossed, from the quadrupedal baboon to the nearly erect gibbon. And in seeking to follow this development through its successive stages, the first point to be considered is how the apes gained their special power of grasping, that characteristic to which they undoubtedly owe the partial freedom of their hands and their tendency to assume the erect attitude.
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