Part 13

Biology needs no apologist when she demands a place--and a prominent place--in any scheme of education worthy of the name. Leave out the Physiological sciences from your curriculum, and you launch the student into the world, undisciplined in that science whose subject-matter would best develop his powers of observation; ignorant of facts of the deepest importance for his own and others' welfare; blind to the richest sources of beauty in God's creation; and unprovided with that belief in a living law, and an order manifesting itself in and through endless change and variety, which might serve to check and moderate that phase of despair through which, if he take an earnest interest in social problems, he will assuredly sooner or later pass.

Finally, one word for myself. I have not hesitated to speak strongly where I have felt strongly; and I am but too conscious that the indicative and imperative moods have too often taken the place of the more becoming subjunctive and conditional. I feel, therefore, how necessary it is to beg you to forget the personality of him who has thus ventured to address you, and to consider only the truth or error in what has been said.

FOOTNOTES:

"In the third place, we have to review the method of Comparison, which is so specially adapted to the study of living bodies, and by which, above all others, that study must be advanced. In Astronomy, this method is necessarily inapplicable; and it is not till we arrive at Chemistry that this third means of investigation can be used, and then only in subordination to the two others. It is in the study, both statical and dynamical, of living bodies that it first acquires its full development; and its use elsewhere can be only through its application here."--COMTE'S Positive Philosophy, translated by Miss Martineau. Vol. i. p. 372.

By what method does M. Comte suppose that the equality or inequality of forces and quantities and the dissimilarity or similarity of forms--points of some slight importance not only in Astronomy and Physics, but even in Mathematics--are ascertained, if not by Comparison?

"Proceeding to the second class of means,--Experiment cannot but be less and less decisive, in proportion to the complexity of the phaenomena to be explored; and therefore we saw this resource to be less effectual in chemistry than in physics: and we now find that it is eminently useful in chemistry in comparison with physiology. In fact, the nature of the phaenomena seems to offer almost insurmountable impediments to any extensive and prolific application of such a procedure in biology."--Comte, vol i. p. 367.

M. Comte, as his manner is, contradicts himself two pages further on, but that will hardly relieve him from the responsibility of such a paragraph as the above.

"Nouvelle Fonction du Foie considere comme organe producteur de matiere sucree chez l'Homme et les Animaux," par M. Claude Bernard.

"Natural Groups given by Type, not by Definition.... The class is steadily fixed, though not precisely limited; it is given, though not circumscribed; it is determined, not by a boundary-line without, but by a central point within; not by what it strictly excludes, but what it eminently includes; by an example, not by a precept; in short, instead of Definition we have a Type for our director. A type is an example of any class, for instance, a species of a genus, which is considered as eminently possessing the characters of the class. All the species which have a greater affinity with this type-species than with any others, form the genus, and are ranged about it, deviating from it in various directions and different degrees."--WHEWELL, The Philosophy of the Inductive Sciences, vol. i. pp. 476, 477.

Save for the pleasure of doing so, I need hardly point out my obligations to Mr. J.S. Mill's "System of Logic," in this view of scientific method.

VI.

ON THE STUDY OF ZOOLOGY.

Natural history is the name familiarly applied to the study of the properties of such natural bodies as minerals, plants, and animals; the sciences which embody the knowledge man has acquired upon these subjects are commonly termed Natural Sciences, in contradistinction to other, so-called "physical," sciences; and those who devote themselves especially to the pursuit of such sciences have been, and are, commonly termed "Naturalists."

Linnaeus was a naturalist in this wide sense, and his "Systema Naturae" was a work upon natural history, in the broadest acceptation of the term; in it, that great methodizing spirit embodied all that was known in his time of the distinctive characters of minerals, animals, and plants. But the enormous stimulus which Linnaeus gave to the investigation of nature soon rendered it impossible that any one man should write another "Systema Naturae," and extremely difficult for any one to become a naturalist such as Linnaeus was.

Great as have been the advances made by all the three branches of science, of old included under the title of natural history, there can be no doubt that zoology and botany have grown in an enormously greater ratio than mineralogy; and hence, as I suppose, the name of "natural history" has gradually become more and more definitely attached to these prominent divisions of the subject, and by "naturalist" people have meant more and more distinctly to imply a student of the structure and functions of living beings.

However this may be, it is certain that the advance of knowledge has gradually widened the distance between mineralogy and its old associates, while it has drawn zoology and botany closer together; so that of late years it has been found convenient (and indeed necessary) to associate the sciences which deal with vitality and all its phenomena under the common head of "biology;" and the biologists have come to repudiate any blood-relationship with their foster-brothers, the mineralogists.

Certain broad laws have a general application throughout both the animal and the vegetable worlds, but the ground common to these kingdoms of nature is not of very wide extent, and the multiplicity of details is so great, that the student of living beings finds himself obliged to devote his attention exclusively either to the one or the other. If he elects to study plants, under any aspect, we know at once what to call him; he is a botanist, and his science is botany. But if the investigation of animal life be his choice, the name generally applied to him will vary, according to the kind of animals he studies, or the particular phenomena of animal life to which he confines his attention. If the study of man is his object, he is called an anatomist, or a physiologist, or an ethnologist; but if he dissects animals, or examines into the mode in which their functions are performed, he is a comparative anatomist or comparative physiologist. If he turns his attention to fossil animals, he is a palaeontologist. If his mind is more particularly directed to the description, specific discrimination, classification, and distribution of animals, he is termed a zoologist.

For the purposes of the present discourse, however, I shall recognise none of these titles save the last, which I shall employ as the equivalent of botanist, and I shall use the term zoology as denoting the whole doctrine of animal life, in contradistinction to botany, which signifies the whole doctrine of vegetable life.

Employed in this sense, zoology, like botany, is divisible into three great but subordinate sciences, morphology, physiology, and distribution, each of which may, to a very great extent, be studied independently of the other.

Zoological morphology is the doctrine of animal form or structure. Anatomy is one of its branches, development is another; while classification is the expression of the relations which different animals bear to one another, in respect of their anatomy and their development.

Zoological distribution is the study of animals in relation to the terrestrial conditions which obtain now, or have obtained at any previous epoch of the earth's history.

Zoological physiology, lastly, is the doctrine of the functions or actions of animals. It regards animal bodies as machines impelled by certain forces, and performing an amount of work, which can be expressed in terms of the ordinary forces of nature. The final object of physiology is to deduce the facts of morphology, on the one hand, and those of distribution on the other, from the laws of the molecular forces of matter.

Such is the scope of zoology. But if I were to content myself with the enunciation of these dry definitions, I should ill exemplify that method of teaching this branch of physical science, which it is my chief business to-night to recommend. Let us turn away then from abstract definitions. Let us take some concrete living thing, some animal, the commoner the better, and let us see how the application of common sense and common logic to the obvious facts it presents, inevitably leads us into all these branches of zoological science.

I have before me a lobster. When I examine it, what appears to be the most striking character it presents? Why, I observe that this part which we call the tail of the lobster, is made up of six distinct hard rings and a seventh terminal piece. If I separate one of the middle rings, say the third, I find it carries upon its under surface a pair of limbs or appendages, each of which consists of a stalk and two terminal pieces. So that I can represent a transverse section of the ring and its appendages upon the diagram board in this way.

If I now take the fourth ring I find it has the same structure, and so have the fifth and the second; so that, in each of these divisions of the tail, I find parts which correspond with one another, a ring and two appendages; and in each appendage a stalk and two end pieces. These corresponding parts are called, in the technical language of anatomy, "homologous parts." The ring of the third division is the "homologue" of the ring of the fifth, the appendage of the former is the homologue of the appendage of the latter. And, as each division exhibits corresponding parts in corresponding places, we say that all the divisions are constructed upon the same plan. But now let us consider the sixth division. It is similar to, and yet different from, the others. The ring is essentially the same as in the other divisions; but the appendages look at first as if they were very different; and yet when we regard them closely, what do we find? A stalk and two terminal divisions, exactly as in the others, but the stalk is very short and very thick, the terminal divisions are very broad and flat, and one of them is divided into two pieces.

I may say, therefore, that the sixth segment is like the others in plan, but that it is modified in its details.

The first segment is like the others, so far as its ring is concerned, and though its appendages differ from any of those yet examined in the simplicity of their structure, parts corresponding with the stem and one of the divisions of the appendages of the other segments can be readily discerned in them.

Thus it appears that the lobster's tail is composed of a series of segments which are fundamentally similar, though each presents peculiar modifications of the plan common to all. But when I turn to the fore part of the body I see, at first, nothing but a great shield-like shell, called technically the "carapace," ending in front in a sharp spine, on either side of which are the curious compound eyes, set upon the ends of stout moveable stalks. Behind these, on the under side of the body, are two pairs of long feelers, or antennae, followed by six pairs of jaws, folded against one another over the mouth, and five pairs of legs, the foremost of these being the great pinchers, or claws, of the lobster.