Part 13

By this process of analysis we reach in our animal or plant those peculiarities which are common to the whole animal or vegetable kingdom, and then we have exhausted the structure so completely that we have nothing remaining to take into account beyond the cell-structure or homogeneous protoplasm by which we know that it is organic, and not a mineral.

The history of the origin of a type, as species, genus, order, etc., is simply the history of the origin of the structure or structures which define those groups respectively. It is nothing more nor less than this, whether a man or an insect be the object of investigation.

EVIDENCES OF DERIVATION.

α. Of Specific Characters.

The evidences of derivation of species from species, within the limits of the genus, are abundant and conclusive. In the first place, the rule which naturalists observe in defining species is a clear consequence of such a state of things. It is not amount and degree of difference that determine the definition of species from species, but it is the permanency of the characters in all cases and under all circumstances. Many species of the systems include varieties and extremes of form, etc., which, were they at all times distinct, and not connected by intermediate forms, would be estimated as species by the same and other writers, as can be easily seen by reference to their works.

Thus, species are either “restricted” or “protean,” the latter embracing many, the former few variations; and the varieties included by the protean species are often as different from each other in their typical forms as are the “restricted” species. As an example, the species Homo sapiens (man) will suffice. His primary varieties are as distinct as the species of many well-known genera, but cannot be defined, owing to the existence of innumerable intermediate forms between them.

As to the common origin of such “varieties” of the protean species, naturalists never had any doubt, yet when it comes to the restricted “species,” the anti-developmentalist denies it in toto. Thus the varieties of most of the domesticated animals are some of them known—others held with great probability to have had a common origin. Varieties of plumage in fowls and canaries are of every-day occurrence, and are produced under our eyes. The cart-horse and racer, the Shetland pony and the Norman, are without doubt derived from the same parentage. The varieties of pigeons and ducks are of the same kind, but not every one is aware of the extent and amount of such variations. The varieties in many characters seen in hogs and cattle, especially when examples from distant countries are compared, are very striking, and are confessedly equal in degree to those found to define species in a state of nature: here, however, they are not definitive.

It is easy to see that all that is necessary to produce in the mind of the anti-developmentalist the illusion of distinct origin by creation of many of these forms, would be to destroy a number of the intermediate conditions of specific form and structure, and thus to leave remaining definable groups of individuals, and therefore “species.”

That such destructions and extinctions have been going on ever since the existence of life on the globe is well known. That it should affect intermediate forms, such as bind together the types of a protean species as well as restricted species, is equally certain. That its result has been to produce definable species cannot be denied, especially in consideration of the following facts: Protean species nearly always have a wide geographical distribution. They exist under more varied circumstances than do individuals of a more restricted species. The subordinate variations of the protean species are generally, like the restricted species, confined to distinct subdivisions of the geographical area which the whole occupies. As in geological time changes of level have separated areas once continuous by bodies of water or high mountain ranges, so have vast numbers of individuals occupying such areas been destroyed. Important alterations of temperature, or great changes in abundance or character of vegetable life over given areas, would produce the same result.

This part of the subject might be prolonged, were it necessary, but it has been ably discussed by Darwin. The rationale of the “origin of species” as stated by him may be examined a few pages farther on.

β. Of the Characters of Higher Groups.

a. Relations of Structures. The evidences of derivative origin of the structures defining the groups called genera, and all those of higher grade, are of a very different character from those discussed in relation to specific characters; they are more difficult of observation and explanation.

Firstly: It would appear to be supposed by many that the creation of organic types was an irregular and capricious process, variously pursued by its Author as regards time and place, and without definite final aim; and this notwithstanding the wonderful evidences we possess, in the facts of astronomy, chemistry, sound, etc., of His adhesion to harmonious and symmetrical sequences in His modes and plans.

Such regularity of plan is found to exist in the relations of the great divisions of the animal and vegetable kingdoms as at present existing on the earth. Thus, with animals we have a great class of species which consists of nothing more than masses or cells of protoplasmic matter, without distinct organs; or the Protozoa. We have then the Cœlenterata (example, corals,) where the organism is composed of many cells arranged in distinct parts, but where a single very simple system of organs, forming the only internal cavity of the body, does the work of the many systems of the more complex animals. Next, the Echinodermata (such as star-fish) present us with a body containing distinct systems of organs enclosed in a visceral cavity, including a rudimental nervous system in the form of a ring. In the Molluscs to this condition is added additional complication, including extensions of the nervous system from the ring as a starting-point, and a special organ for a heart. In the Articulates (crabs, insects,) we have like complications, and a long distinct nervous axis on the lower surface of the body. The last branch or division of animals is considered to be higher, because all the systems of life organs are most complex or specialized. The nervous ring is almost obliterated by a great enlargement of its usual ganglia, thus become a brain, which is succeeded by a long axis on the upper side of the body. This and other points define the Vertebrata.

Plans of structure, independent of the simplicity or perfection of the special arrangement or structure of organs, also define these great groups. Thus the Protozoa present a spiral, the Cœlenterata a radiate, the Echinodermata a bilateral radiate plan. The Articulates are a series of external rings, each in one or more respects repeating the others. The Molluscs are a sac, while a ring above a ring, joined together by a solid center-piece, represents the plan of each of the many segments of the Vertebrates which give the members of that branch their form.

These bulwarks of distinction of animal types are entered into here simply because they are the most inviolable and radical of those with which we have to deal, and to give the anti-developmentalist the best foothold for his position. I will only allude to the relations of their points of approach, as these are affected by considerations afterward introduced.

The Vertebrates approach the Molluscs at the lowest extreme of the former and higher of the latter. The lamprey eels of the one possess several characters in common with the cuttle-fish or squids of the latter. The amphioxus is called the lowest Vertebrate, and though it is nothing else, the definition of the division must be altered to receive it; it has no brain!

The lowest forms of the Molluscs and Articulates are scarcely distinguishable from each other, so far as adhesion to the “plan” is concerned, and some of the latter division are very near certain Echinodermata. As we approach the boundary-lines of the two lowest divisions, the approaches become equally close, and the boundaries very obscure.

More instructive is the evidence of the relation of the subordinate classes of any one of these divisions. The conditions of those organs or parts which define classes exhibit a regular relation, commencing with simplicity and ending with complication; first associated with weak exhibitions of the highest functions of the nervous system—at the last displaying the most exalted traits found in the series.

For example: In the classes of Vertebrates we find the lowest nervous system presents great simplicity—the brain cannot be recognized; next (in lampreys), the end of the nervous axis is subdivided, but scarcely according to the complex type that follows. In fishes the cerebellum and cerebral hemispheres are minute, and the intermediate or optic lobes very large: in the reptiles the cerebral hemispheres exceed the optic lobes, while the cerebellum is smaller. In birds the cerebellum becomes complex and the cerebrum greatly increases. In mammals the cerebellum increases in complexity or number of parts, the optic lobes diminish, while the cerebral hemispheres become wonderfully complex and enlarged, bringing us to the highest development, in man.

The history of the circulatory system in the Vertebrates is the same. First, a heart with one chamber, then one with two divisions: three divisions belong to a large series, and the highest possess four. The origins of the great artery of the body, the aorta, are first five on each side: they lose one in the succeeding class in the ascending scale, and one in each succeeding class or order, till the Mammalia, including man, present us with but one on one side.

Footnote 45:

See a homological system of the circulatory system in the author’s Origin of Genera, p. 22.

From an infinitude of such considerations as the above, we derive the certainty that the general arrangement of the various groups of the organic world is in scales, the subordinate within the more comprehensive divisions. The identification of all the parts in such a complexity of organism as the highest animals present, is a matter requiring much care and attention, and constitutes the study of homologies. Its pursuit has resulted in the demonstration that every individual of every species of a given branch of the animal kingdom is composed of elements common to all, and that the differences which are so radical in the higher groups are but the modifications of the same elemental parts, representing completeness or incompleteness, obliteration or subdivision. Of the former character are rudimental organs, of which almost every species possesses an example in some part of its structure.

But we have other and still more satisfactory evidence of the meaning of these relations. By the study of embryology we can prove most indubitably that the simple and less complex are inferior to the more complex. Selecting the Vertebrates again as an example, the highest form of mammal—e.g., man—presents in his earliest stages of embryonic growth a skeleton of cartilage, like that of the lamprey: he also possesses five origins of the aorta and five slits on the neck, both which characters belong to the lamprey and the shark. If the whole number of these parts does not coexist in the embryonic man, we find in embryos of lower forms more nearly related to the lamprey that they do. Later in the life of the mammal but four aortic origins are found, which arrangement, with the heart now divided into two chambers, from a beginning as a simple tube, is characteristic of the class of Vertebrates next in order—the bony fishes. The optic lobes of the human brain have also at this time a great predominance in size—a character above stated to be that of the same class. With advancing development the infant mammal follows the scale already pointed out. Three chambers of the heart and three aortic origins follow, presenting the condition permanent in the batrachia; and two origins, with enlarged cerebral hemispheres of the brain, resemble the reptilian condition. Four heart-chambers, and one aortic root on each side, with slight development of the cerebellum, follow all characters defining the crocodiles, and immediately precede the special conditions defining the mammals. These are, the single aorta root from one side, and the full development of the cerebellum: later comes that of the cerebrum also in its higher mammalian and human traits.

Thus we see the order already pointed out to be true, and to be an ascending one. This is the more evident as each type or class passes through the conditions of those below it, as did the mammal; each scale being shorter as its highest terminus is lower. Thus the crocodile passes through the stage of the lamprey, the fish, the batrachian and the reptile proper.

b. In Time. We have thus a scale of relations of existing forms of animals and plants of a remarkable kind, and such as to stimulate greatly our inquiries as to its significance. When we turn to the remains of the past creation preserved to us in the deposits continued throughout geologic time, we are not disappointed, for great light is at once thrown upon the subject.

We find, in brief, that the lowest division of the animal kingdom appeared first, and long before any type of a higher character was created. The Protozoön, Eozoön, is the earliest of animals in geologic time, and represents the lowest type of animal life now existing. We learn also that the highest branch appeared last. No remains of Vertebrates have been found below the lower Devonian period, or not until the Echinoderms and Molluscs had reached a great preëminence. It is difficult to be sure whether the Protozoa had a greater numerical extent in the earliest periods than now, but there can be no doubt that the Cœlenterata (corals) and Echinoderms (crinoids) greatly exceeded their present bounds, in Paleozoic time, so that those at present existing are but a feeble remnant. If we examine the subdivisions known as classes, evidence of the nature of the succession of creation is still more conclusive. The most polyp-like of the Molluscs (brachiopoda) constituted the great mass of its representatives during Paleozoic time. Among Vertebrates the fishes appear first, and had their greatest development in size and numbers during the earliest periods of the existence of the division. Batrachia were much the largest and most important of land animals during the Carboniferous period, while the higher Vertebrates were unknown. The later Mesozoic periods saw the reign of reptiles, whose position in structural development has been already stated. Finally, the most perfect, the mammal, came upon the scene, and in his humblest representatives. In Tertiary times mammalia supplanted the reptiles entirely, and the unspiritual mammals now yield to man, the only one of his class in whom the Divine image appears.