Part 6

Specimens examined.--Total 7. Fremont Co.: Garden Park, Canyon City, 5344 ft., 1; 2-9/10 mi. E Canyon City, 5344 ft., 6.

SUMMARY

A study of 249 specimens of Thomomys bottae from Colorado reveals six subspecies in the state. T. b. aureus and T. b. howelli occupy the Colorado Plateau Region in the western and southwestern parts of the state. T. b. internatus, T. b. cultellus, T. b. pervagus, and the newly named T. b. rubidus occupy part of the Southern Rocky Mountain Region and a narrow strip of the Great Plains.

The greatest amount of geographic variation, in Thomomys bottae in Colorado, occurs in the ecotone between the grassland and coniferous forest at the edge of the Great Plains, and in the ecotone between the Pinon, juniper, and sage of the Colorado Plateau and the Coniferous forest of the southern Rocky mountains.

TABLE 1. MEASUREMENTS, IN MILLIMETERS, OF THOMOMYS BOTTAE

Unless otherwise noted, specimens are adults from Colorado

Key to Headings: A: Catalog number or number of individuals averaged B: Total length C: Tail D: Hind foot E: Condylobasal length F: Nasal length G: Zygomatic breadth H: Squamosal breadth I: Length of rostrum J: Breadth of rostrum K: Alveolar length of upper max. tooth-row L: Least interorbital breadth

=============================================================== Sex| |||| | | | | | || ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | Thomomys bottae howelli, holotype ---+---------+---+---+---+----+----+----+----+----+----+---+--- F | 75684| | | | | | | | | | | | sad. |219| 71| 29|37.3|11.1|23.7|20.0|14.5| 8.5|7.7|6.6 | +---+---+---+----+----+----+----+----+----+---+--- | | Thomomys bottae aureus, Bedrock | +---+---+---+----+----+----+----+----+----+---+--- F | 2982 |217| 59| 31|40.4|13.8|24.3|20.6|16.7| 8.6|9.2|6.8 F | 3013 |210| 60| 29|38.7|13.0|24.4|20.4|15.7| 8.1|8.9|7.0 | | | | | | | | | | | | M | 2997 |242| 73| 33|44.7|15.4|28.4|22.8|15.7|10.1|9.0|7.2 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | Coventry | +---+---+---+----+----+----+----+----+----+---+--- F | 6 av. |222| 61| 31|39.0|12.1|25.4|20.3|15.2| 7.8|8.2|6.8 | Max. |229| 63| 33|40.0|12.8|25.8|20.6|15.6| 8.2|8.5|7.0 | Min. |217| 58| 30|38.3|11.4|25.0|19.3|14.7| 7.5|8.0|6.6 | | | | | | | | | | | | M | 3 av. |259| 70| 35|46.5|15.0|29.3|22.7|17.8| 9.2|9 5|6.7 | Max. |270| 76| 36|48.3|16.0|31.9|23.7|18.0| 9.3|9.6|6.9 | Min. |250| 65| 35|45.5|14.4|27.6|22.2|17.6| 9.2|8.0|6.4 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | Ashbaugh's Ranch and 15 mi. W Cortez | +---+---+---+----+----+----+----+----+----+---+--- F | 7 av. |225| 67| 28|39.0|13.7|24.4|20.2|16.1| 8.0|8.3|6.3 | Max. |238| 75| 31|40.6|14.7|25.0|20.7|16.5| 8.5|8.7|6.8 | Min. |216| 55| 26|37.8|12.9|23.6|19.7|15.5| 7.8|7.9|6.1 | | | | | | | | | | | | M | 4 av. |247| 73| 31|44.2|15.9|27.7|22.1|18.6| 9.2|8.4|6.4 | Max. |252| 80| 34|45.2|16.7|28.8|22.3|19.8| 9.6|8.8|6.7 | Min. |244| 67| 30|43.7|15.5|27.0|21.7|18.0| 8.8|8.0|6.2 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | Cortez | +---+---+---+----+----+----+----+----+----+---+--- F | 5120 |224| 56| 28|38.1|12.3|....|19.5|15.4| 7.5|7.6|6.5 | 5121 |220| 68| 31|38.3|11.6|24.2|19.6|15.1| 7.6|8.0|6.7 | | | | | | | | | | | | M | 5124 |257| 81| 33|44.4|15.4|29.5|22.2|18.6| 8.9|8.6|6.5 M | 5119 |215| 62| 28|42.0|14.0|27.9|22.1|17.9| 8.2|8.6|6.4 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | Mesa Verde (combined) | +---+---+---+----+----+----+----+----+----+---+--- F | 5 av. |221| 63| 30|39.0|12.6|24.7|20.2|16.0| 8.0|8.2|6.7 | Max. |235| 66| 32|40.4|13.7|25.6|21.1|17.3| 8.7|8.5|7.1 | Min. |212| 61| 28|38.1|12.0|24.1|19.5|15.0| 7.7|7.9|6.4 | | | | | | | | | | | | M | 3 av. |246| 74| 32|43.7|14.9|27.8|22.3|18.3| 8.8|8.9|6.6 | Max. |252| 79| 33|45.0|15.2|28.4|23.0|18.5| 9.0|9.0|6.8 | Min. |238| 69| 31|42.0|14.7|27.5|21.2|18.2| 8.7|8.9|6.3 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | 1 mi. N La Plata | +---+---+---+----+----+----+----+----+----+---+--- M | 72966|236| 70| 31|45.4|15.6|29.4|23.3|20.2| 8.8|8.5|6.5 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | 3 mi. W Durango | +---+---+---+----+----+----+----+----+----+---+--- F | 3 av. |225| 65| 28|40.1|13.1|25.7|21.0|16.6| 8.1|8.3|6.5 | Max. |230| 67| 29|40.4|13.5|25.8|21.2|16.8| 8.4|8.5|6.6 | Min. |219| 63| 28|39.9|13.0|25.7|20.6|16.5| 8.4|8.1|6.4 | | | | | | | | | | | | M | 70054|262| 87| 35|45.0|15.6|27.9|22.7|19.7| 9.3|9.8|6.4 M | 70055|248| 79| 31|43.3|14.0|27.6|22.1|17.1| 8.7|8.2|6.2 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | 12 mi. W Pagosa Springs | +---+---+---+----+----+----+----+----+----+---+--- F | 72971|217| 65| 27|39.1|12.8|....|20.0|15.4| 7.4|8.7|6.2 | | | | | | | | | | | | M | 72970|238| 70| 29|42.7|15.0|27.5|21.8|17.2| 8.8|8.3|6.5 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | Thomomys bottae pervagus, Antonito | +---+---+---+----+----+----+----+----+----+---+--- F |133668| | | | | | | | | | | | sad. |208| 69| 29|37.3|12.9|23.1|18.2|15.8| 7.5|8.0|6.9 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | Espanola, New Mexico | +---+---+---+----+----+----+----+----+----+---+--- F |133616|249| 82| 38|41.1|....|24.6|20.0|16.3| 8.2|8.1|7.1 F |133619|216| 65| 32|40.6|....|24.9|19.3|....| 8.0|8.0|6.8 | | | | | | | | | | | | M | 58293|244| 76| 31|44.0|16.1|26.9|21.2|18.3| 8.8|8.1|6.6 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | Thomomys bottae internatus, Salida | +---+---+---+----+----+----+----+----+----+---+--- F | 11 av. |219| 67| 31|38.6|13.4|23.2|19.5|15.4| 7.6|7.8|6.5 | Max. |242| 80| 34|40.4|14.2|25.0|20.2|16.2| 8.1|8.4|6.9 | Min. |196| 45| 29|37.6|12.9|21.9|18.8|14.8| 7.3|7.0|6.3 | | | | | | | | | | | | M | 3 av. |247| 74| 32|42.9|16.1|25.1|20.9|18.0| 8.2|8.0|6.3 | Max. |248| 74| 33|43.7|16.3|26.4|21.7|18.1| 8.8|8.1|6.4 | Min. |247| 74| 32|42.2|15.9|25.8|20.5|17.9| 7.9|7.9|6.3 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | 12 mi. N Canyon City | +---+---+---+----+----+----+----+----+----+---+--- F | 72945|230| 81| 28|38.1|13.0|22.6|19.4|15.0| 7.9|8.0|6.7 F | 72947|228| 74| 27|38.7|14.0|23.6|19.8|15.9| 8.2|8.1|6.8 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | 1 mi. W Coaldale | +---+---+---+----+----+----+----+----+----+---+--- F | 70042|224| 70| 30|38.1|13.1|23.5|19.5|15.6| 7.7|7.5|6.6 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | 5 mi. S Cotopaxi | +---+---+---+----+----+----+----+----+----+---+--- F | 72932|224| 65| 27|39.1|13.8|24.3|20.4|15.5| 7.7|7.5|6.5 | | | | | | | | | | | | M | 72925|250| 74| 29|44.0|16.2|27.5|22.8|18.7| 9.0|8.1|6.1 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | 9 mi. SSW Colorado Springs | +---+---+---+----+----+----+----+----+----+---+--- F | 72942|225| 77| 29|38.8|14.1|23.3|20.2|15.4| 7.8|8.3|6.7 | 72943|219| 70| 28|37.7|13.5|23.0|19.7|14.8| 7.6|8.4|6.8 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | 2-1/2 mi. S Wetmore | +---+---+---+----+----+----+----+----+----+---+--- M | 70053|250| 81| 30|42.5|16.7|26.3|22.3|17.7| 8.5|7.9|5.9 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | 200 yards E St. Charles River, 8 mi. W Pueblo | +---+---+---+----+----+----+----+----+----+---+--- F | 73497|226| 69| 30|39.3|13.9|24.9|20.5|15.7| 7.7|7.9|7.2 F | 73498|216| 64| 29|38.0|12.9|24.2|20.1|15.1| 7.7|7.4|6.7 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | St. Charles Mesa | +---+---+---+----+----+----+----+----+----+---+--- F | 4860 |222| 70| 29|38.2|13.5|....|19.3|15.9| 8.2|7.5|6.5 M | 4864 |240| 72| 33|43.1|15.8|....|21.4|17.6| 9.2|7.9|6.7 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | 11 mi. WNW Gardner | +---+---+---+----+----+----+----+----+----+---+--- F | 70052|227| 64| 28|37.9|13.0|22.5|18.8|14.8| 7.3|8.0|6.7 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | 1-1/2 mi. S Redwing | +---+---+---+----+----+----+----+----+----+---+--- F | 72940|227| 73| 28|39.0|13.1|23.1|18.8|15.6| 7.8|8.0|6.8 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | 1 mi. E La Veta | +---+---+---+----+----+----+----+----+----+---+--- M | 70049|254| 88| 32|42.4|15.1|27.5|21.8|17.3| 8.4|8.2|6.5 M | 70044|239| 80| 32|42.3|16.5|27.8|22.0|17.9| 8.7|8.1|6.4 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | Thomomys bottae cultellus, Fishers Peak | +---+---+---+----+----+----+----+----+----+---+--- F |129285| | | | | | | | | | | | sad. |214| 64| 27|37.2|13.0|....|19.0|15.3| 7.7|7.6|6.5 ---+---------+---+---+---+----+----+----+----+----+----+---+--- | | Thomomys bottae rubidus, holotype and topotypes | +---+---+---+----+----+----+----+----+----+---+--- F | 72952|233| 80| 28|40.6|14.2|25.1|20.8|16.7| 8.8|7.5|6.9 F | 72954|225| 80| 28|40.3|14.2|24.6|20.6|16.6| 9.2|7.2|6.9 | | | | | | | | | | | | M | 3 av. |261| 89| 31|44.7|15.7|27.8|22.6|18.6|10.1|7.4|6.9 | Max. |270| 94| 32|45.1|15.9|28.1|22.7|18.8|10.4|7.6|7.0 | Min. |255| 85| 30|44.2|15.5|27.5|22.5|18.5| 9.8|7.2|6.8 ---+---------+---+---+---+----+----+----+----+----+----+---+---

sad. denotes subadult. 1. United States National Museum. 2. E. R. Warren Collection. 3. Colorado State University. 4. Museum of Natural History, University of Kansas.

LITERATURE CITED

BAILEY, V.

1910. Two new pocket gophers of the genus Thomomys. Proc. Biol. Soc. Washington, 23:79-80, May 4.

1915. Revision of the pocket gophers of the genus Thomomys. U. S. Dept. Agric., Bur. Biol. Surv., N. Amer. Fauna, 39:1-136, 8 pls., 10 figs, in text, November 15.

DAUBENMIRE, R. F.

1943. Vegetational zonation in the Rocky Mountains. Bot. Rev., 9:325-393, June.

DURRANT, S. D.

1952. Mammals of Utah. Univ. Kansas Publ., Mus. Nat. Hist., 6:1-549, 91 figs. August 10.

FENNEMAN, N. M.

1931. Physiography of western United States. McGraw Hill Book Co., New York, xiii + 534 pp., 173 figs., 1 map in cover pocket.

GOLDMAN, E. A.

1936. New pocket gophers of the genus Thomomys. Jour. Washington Acad. Sci., 26(3):111-120, March 15.

GRINNELL, J.

1931. A new pocket gopher from southeastern California. Univ. California Publ. Zool., 38(1):1-10, 2 pls., October 17.

KELSON, K. R.

1951. Two new subspecies of Thomomys bottae from New Mexico and Colorado. Univ. Kansas Publ., Mus. Nat. Hist, 5(6):59-71, 1 fig. in text, October 1.

MUNSELL, A. H.

1954. Munsell soil color charts. Munsell Color Co., Inc., Baltimore.

Transmitted November 14, 1957.