Part 3

Additional, marginal records (Durrant, Univ. Kansas Publ., Mus. Nat. Hist., 6:356, August 19, 1952).--+Utah+: Rich Co.: Monte Cristo, 18 mi. W Woodruff, 8000 ft. Salt Lake Co.: Emigration Canyon, 8 mi. above forks, 6,000 ft.; Silver Lake Post Office (Brighton), 9,500 ft. Wasatch Co.: Wolf Creek Summit, 9,800 ft. Daggett Co.: Beaver Dams, 10,500 ft.

+Clethrionomys gapperi gauti+, new subspecies

Type.--Male, adult, skin and skull; No. 133515, Biological Surveys Collections, United States National Museum, from Twining, 10,700 ft., Taos County, New Mexico; obtained on August 7, 1904, by James H. Gaut, original number 3086.

Range.--The Rocky Mountains of north-central New Mexico and south-central Colorado.

Diagnosis.--A brightly colored Clethrionomys gapperi; dorsal stripe near Chestnut (capitalized color terms after Ridgway, Color Standards and Color Nomenclature, Washington, D. C., 1912) with an underwash of near Light Ochraceous-Buff and a mixture of black-tipped guard hairs giving an over-all effect of between Tawny and Russet; braincase relatively large; zygomatic width and lambdoidal width large; nasals long.

Comparisons.--As compared with topotypes of C. g. galei, the color is lighter, the dorsal reddish stripe slightly narrower, sides brighter, with a wash of Light Ochraceous-Buff, grading ventrally into a slight wash of Pale Ochraceous-Buff, instead of a silvery-white venter characteristic of C. g. galei; zygomatic and lambdoidal breadths are greater, nasals slightly shorter, auditory bullae slightly more inflated, teeth larger, and braincase larger.

As compared with topotypes of C. g. limitis, C. g. gauti is darker, has a greater zygomatic breadth, longer upper tooth-row, longer nasals, and narrower rostrum.

Measurements.--External and cranial measurements of the type, and the average and extreme measurements of four adult males and one adult female from the type locality (including the type) and five miles south of the type locality are: Total length, 144, 147 (140-152); tail, 40, 42 (39-45); hind foot, 20, 19.3 (19-20); condylobasilar length, 22.3, 22.9 (22.2-24.0); zygomatic breadth, 13.6, 13.7 (13.5-14.0); lambdoidal breadth, 11.9, 11.7 (11.4-12.0); alveolar length of upper cheek-teeth, 5.1, 5.2 (5.1-5.4); interorbital breadth, 4.0, 3.9 (3.8-4.0); length of nasals, 7.0, 7.2 (7.0-7.6); breadth of rostrum, 2.9, 3.1 (2.9-3.4); length of incisive foramina, 4.8, 5.1 (4.8-5.3).

Remarks.--Two specimens from a locality 21 mi. W and 3 mi. N Saguache, Saguache County, Colorado, although referred to this subspecies on the basis of paler pelage, inflation of auditory bullae, and heavier teeth, show characters of C. g. galei in the narrowness across the zygomata and lambdoidal crest. Four specimens from Silverton (1 adult and 3 young adults) are referable to this subspecies on the basis of color of pelage and cranial proportions but are smaller than either C. g. gauti or C. g. galei.

The specimen from Pecos Baldy, Pecos Mountain, San Miguel County, New Mexico, referred by Bailey (N. Amer. Fauna, 52:192) to Clethrionomys gapperi galei, is here referred to C. g. gauti on geographical grounds.

The name gauti is proposed in honor of the collector of the type specimen, James H. Gaut.

Specimens examined.--Total, 14, distributed as follows and, unless otherwise stated, in the Biological Surveys Collection:

+Colorado+: Saguache County: 21 mi. W and 3 mi. N Saguache, N 38°, 106° 31´, 9100 ft., 2 (Univ. Kansas Mus. Nat. Hist.). San Juan Co.: Silverton, 4.

+New Mexico+: Taos County: Twining, 10,700 ft., 3; 5 mi. S Twining, 11,400 ft., 3. Sandoval County: Goat Peak, Jemez Mountains, 1. Colfax County: 15 mi. SW Cimarron, 9000 ft., 1 (Amer. Mus. Nat. Hist.).

Additional records.--+New Mexico+: San Miguel Co.: Pecos Baldy, Pecos Mountain, 1 (Bailey, N. Amer. Fauna, 52:192, 1932).

+Clethrionomys gapperi limitis+ (Bailey)

1913. Evotomys limitis Bailey, Proc. Biol. Soc. Washington, 26:133, May 21.

Type locality.--Willow Creek, a branch of the Gilita, 8500 ft., Mogollon Mountains, Catron County, New Mexico.

Range.--Known from the Mogollon, San Mateo, and Magdalena mountains of western New Mexico.

Remarks.--Bailey (Proc. Biol. Soc. Washington, 26:133, May 21, 1913) described this animal as a species and gave as general characteristics: "Size slightly larger than E. [= Clethrionomys gapperi] galei; colors duller, grayer and less buffy; skull and dentition heavier." He further characterized the skull as: "Larger, heavier and conspicuously more ridged than in galei; bullae large and especially deep; dentition heavy throughout." The type of C. limitis, as judged from the measurements given by Bailey (loc. cit.), is an exceptionally old male.

Our comparison of six adult topotypes with a series of C. g. galei from Wyoming (18 adults from 3 mi. SSE Browns Peak, 10,000 ft., Albany County, in Univ. Kansas Mus. Nat. Hist.) and with three near-topotypes of C. g. galei (3 mi. S Ward, 9000 ft., Boulder County, Colorado, in Univ. Kansas Mus. Nat. Hist.) revealed that most of the differences noted by Bailey (loc. cit.) are not evident when individuals of comparable ages are examined. Some specimens of C. g. galei exceed C. limitis in ridging of the skull and size of the teeth although conspicuous ridges and large teeth are supposedly distinctive of C. limitis. The bullae, although averaging larger in C. limitis, can be matched in size by those of specimens of C. g. galei from Wyoming.

The differences evident between C. limitis and C. g. galei are of the kind and degree that serve to separate subspecies in the species Clethrionomys gapperi and, although actual evidence of intergradation is lacking, we think that the relationships of limitis are better expressed by arranging it as a subspecies of C. gapperi than by retaining it as a full species.

Specimens examined.--Total, 7, all in the Biological Surveys Collection, distributed as follows:

+New Mexico+: Catron County: Willow Creek, 8500 ft., Mogollon Mountains, 4. Socorro County: San Mateo Peak, 10,000 ft., San Mateo Mountains, 2; Copper Canyon, 9000 ft., Magdalena Mountains, 1.

+Clethrionomys gapperi arizonensis+, new subspecies

Type.--Female, adult, skin and skull; No. 158401, Biological Surveys Collection, United States National Museum; from Little Colorado River, 8300 ft., White Mountains, Apache County, Arizona; obtained September 12, 1908, by C. Birdseye, original number 152.

Range.--Known only from the White Mountains of eastern Arizona.

Diagnosis.--Dorsal stripe near Chestnut, with an underwash of between Tawny and Russet, and a mixture of black-tipped hairs, resulting in an overall effect of near Chestnut. Skull wide across zygomatic arches and narrow across mastoids; rostrum narrow and posterior border of palate straight.