It is certainly the Pongo of Wurmb; and it is as certainly not the Pongo of Battell, seeing that the Orang-Utan is entirely confined to the great Asiatic islands of Borneo and Sumatra.
And while the progress of discovery thus cleared up the history of the Orang, it also became established that the only other man-like Apes in the eastern world were the various species of Gibbon--Apes of smaller stature, and therefore attracting less attention than the Orangs, though they are spread over a much wider range of country, and are hence more accessible to observation.
Although the geographical area inhabited by the 'Pongo' and Engeco of Battell is so much nearer to Europe than that in which the Orang and Gibbon are found, our acquaintance with the African Apes has been of slower growth; indeed, it is only within the last few years that the truthful story of the old English adventurer has been rendered fully intelligible. It was not until 1835 that the skeleton of the adult Chimpanzee became known, by the publication of Professor Owen's above-mentioned very excellent memoir 'On the osteology of the Chimpanzee and Orang', in the 'Zoological Transactions'--a memoir which, by the accuracy of its descriptions, the carefulness of its comparisons, and the excellence of its figures, made an epoch in the history of our knowledge of the bony framework, not only of the Chimpanzee, but of all the anthropoid Apes.
By the investigations herein detailed, it became evident that the old Chimpanzee acquired a size and aspect as different from those of the young known to Tyson, to Buffon, and to Traill, as those of the old Orang from the young Orang; and the subsequent very important researches of Messrs. Savage and Wyman, the American missionary and anatomist, have not only confirmed this conclusion, but have added many new details.
One of the most interesting among the many valuable discoveries made by Dr. Thomas Savage is the fact, that the natives in the Gaboon country at the present day, apply to the Chimpanzee a name--"Enche-eko"--which is obviously identical with the "Engeko" of Battell; a discovery which has been confirmed by all later inquirers. Battell's "lesser monster" being thus proved to be a veritable existence, of course a strong presumption arose that his "greater monster," the 'Pongo,' would sooner or later be discovered. And, indeed, a modern traveller, Bowdich, had, in 1819, found strong evidence, among the natives, of the existence of a second great Ape, called the 'Ingena,' "five feet high, and four across the shoulders," the builder of a rude house, on the outside of which it slept.
In 1847, Dr. Savage had the good fortune to make another and most important addition to our knowledge of the man-like Apes; for, being unexpectedly detained at the Gaboon river, he saw in the house of the Rev. Mr. Wilson, a missionary resident there, "a skull represented by the natives to be a monkey-like animal, remarkable for its size, ferocity, and habits." From the contour of the skull, and the information derived from several intelligent natives, "I was induced," says Dr. Savage (using the term Orang in its old general sense) "to believe that it belonged to a new species of Orang. I expressed this opinion to Mr. Wilson, with a desire for further investigation; and, if possible, to decide the point by the inspection of a specimen alive or dead." The result of the combined exertions of Messrs. Savage and Wilson was not only the obtaining of a very full account of the habits of this new creature, but a still more important service to science, the enabling the excellent American anatomist already mentioned, Professor Wyman, to describe, from ample materials, the distinctive osteological characters of the new form. This animal was called by the natives of the Gaboon "Enge-ena," a name obviously identical with the "Ingena" of Bowdich; and Dr. Savage arrived at the conviction that this last discovered of all the great Apes was the long-sought "Pongo" of Battell.
The justice of this conclusion, indeed, is beyond doubt--for not only does the 'Enge-ena' agree with Battell's "greater monster" in its hollow eyes, its great stature, and its dun or iron-grey colour, but the only other man-like Ape which inhabits these latitudes--the Chimpanzee--is at once identified, by its smaller size, as the "lesser monster," and is excluded from any possibility of being the 'Pongo,' by the fact that it is black and not dun, to say nothing of the important circumstance already mentioned that it still retains the name of 'Engeko,' or "Enche-eko," by which Battell knew it.
In seeking for a specific name for the "Enge-ena," however, Dr. Savage wisely avoided the much misused 'Pongo'; but finding in the ancient Periplus of Hanno the word "Gorilla" applied to certain hairy savage people, discovered by the Carthaginian voyager in an island on the African coast, he attached the specific name "Gorilla" to his new ape, whence arises its present well-known appellation. But Dr. Savage, more cautious than some of his successors, by no means identifies his ape with Hanno's "wild men." He merely says that the latter were "probably one of the species of the Orang;" and I quite agree with M. Brulle, that there is no ground for identifying the modern 'Gorilla' with that of the Carthaginian admiral.
Since the memoir of Savage and Wyman was published, the skeleton of the Gorilla has been investigated by Professor Owen and by the late Professor Duvernoy, of the Jardin des Plantes, the latter having further supplied a valuable account of the muscular system and of many of the other soft parts; while African missionaries and travellers have confirmed and expanded the account originally given of the habits of this great man-like Ape, which has had the singular fortune of being the first to be made known to the general world and the last to be scientifically investigated.
Two centuries and a half have passed away since Battell told his stories about the 'greater' and the 'lesser monsters' to Purchas, and it has taken nearly that time to arrive at the clear result that there are four distinct kinds of Anthropoids--in Eastern Asia, the Gibbons and the Orangs; in Western Africa, the Chimpanzees and the Gorilla.
The man-like Apes, the history of whose discovery has just been detailed, have certain characters of structure and of distribution in common. Thus they all have the same number of teeth as man--possessing four incisors, two canines, four false molars, and six true molars in each jaw, or 32 teeth in all, in the adult condition; while the milk dentition consists of 20 teeth--or four incisors, two canines, and four molars in each jaw. They are what are called catarrhine Apes--that is, their nostrils have a narrow partition and look downwards; and, furthermore, their arms are always longer than their legs, the difference being sometimes greater and sometimes less; so that if the four were arranged in the order of the length of their arms in proportion to that of their legs, we should have this series--Orang (1 4/9:1), Gibbon (1 1/4:1), Gorilla (1 1/5:1), Chimpanzee (1 1/16:1). In all, the fore limbs are terminated by hands, provided with longer or shorter thumbs; while the great toe of the foot, always smaller than in Man, is far more movable than in him and can be opposed, like a thumb, to the rest of the foot. None of these apes have tails, and none of them possess the cheek pouches common among monkeys. Finally, they are all inhabitants of the old world.
The Gibbons are the smallest, slenderest, and longest-limbed of the man-like apes: their arms are longer in proportion to their bodies than those of any of the other man-like Apes, so that they can touch the ground when erect; their hands are longer than their feet, and they are the only Anthropoids which possess callosities like the lower monkeys. They are variously coloured. The Orangs have arms which reach to the ankles in the erect position of the animal; their thumbs and great toes are very short, and their feet are longer than their hands. They are covered with reddish brown hair, and the sides of the face, in adult males, are commonly produced into two crescentic, flexible excrescences, like fatty tumours. The Chimpanzees have arms which reach below the knees; they have large thumbs and great toes, their hands are longer than their feet; and their hair is black, while the skin of the face is pale. The Gorilla, lastly, has arms which reach to the middle of the leg, large thumbs and great toes, feet longer than the hands, a black face, and dark-grey or dun hair.
For the purpose which I have at present in view, it is unnecessary that I should enter into any further minutiae respecting the distinctive characters of the genera and species into which these man-like Apes are divided by naturalists. Suffice it to say, that the Orangs and the Gibbons constitute the distinct genera, 'Simia' and 'Hylobates'; while the Chimpanzees and Gorillas are by some regarded simply as distinct species of one genus, 'Troglodytes'; by others as distinct genera--'Troglodytes' being reserved for the Chimpanzees, and 'Gorilla' for the Enge-ena or Pongo.
Sound knowledge respecting the habits and mode of life of the man-like Apes has been even more difficult of attainment than correct information regarding their structure.
Once in a generation, a Wallace may be found physically, mentally, and morally qualified to wander unscathed through the tropical wilds of America and of Asia; to form magnificent collections as he wanders; and withal to think out sagaciously the conclusions suggested by his collections: but, to the ordinary explorer or collector, the dense forests of equatorial Asia and Africa, which constitute the favourite habitation of the Orang, the Chimpanzee, and the Gorilla, present difficulties of no ordinary magnitude: and the man who risks his life by even a short visit to the malarious shores of those regions may well be excused if he shrinks from facing the dangers of the interior; if he contents himself with stimulating the industry of the better seasoned natives, and collecting and collating the more or less mythical reports and traditions with which they are too ready to supply him.
In such a manner most of the earlier accounts of the habits of the man-like Apes originated; and even now a good deal of what passes current must be admitted to have no very safe foundation. The best information we possess is that, based almost wholly on direct European testimony respecting the Gibbons; the next best evidence relates to the Orangs; while our knowledge of the habits of the Chimpanzee and the Gorilla stands much in need of support and enlargement by additional testimony from instructed European eye-witnesses.
It will therefore be convenient in endeavouring to form a notion of what we are justified in believing about these animals, to commence with the best known man-like Apes, the Gibbons and Orangs; and to make use of the perfectly reliable information respecting them as a sort of criterion of the probable truth or falsehood of assertions respecting the others.
Of the GIBBONS, half a dozen species are found scattered over the Asiatic islands, Java, Sumatra, Borneo, and through Malacca, Siam, Arracan, and an uncertain extent of Hindostan, on the main land of Asia. The largest attain a few inches above three feet in height, from the crown to the heel, so that they are shorter than the other man-like Apes; while the slenderness of their bodies renders their mass far smaller in proportion even to this diminished height.
Dr. Salomon Muller, an accomplished Dutch naturalist, who lived for many years in the Eastern Archipelago, and to the results of whose personal experience I shall frequently have occasion to refer, states that the Gibbons are true mountaineers, loving the slopes and edges of the hills, though they rarely ascend beyond the limit of the fig-trees. All day long they haunt the tops of the tall trees; and though, towards evening, they descend in small troops to the open ground, no sooner do they spy a man than they dart up the hill-sides, and disappear in the darker valleys.
All observers testify to the prodigious volume of voice possessed by these animals. According to the writer whom I have just cited, in one of them, the Siamang, "the voice is grave and penetrating, resembling the sounds goek, goek, goek, goek, goek ha ha ha ha haaaaa, and may easily be heard at a distance of half a league." While the cry is being uttered, the great membranous bag under the throat which communicates with the organ of voice, the so-called "laryngeal sac," becomes greatly distended, diminishing again when the creature relapses into silence.
M. Duvaucel, likewise, affirms that the cry of the Siamang may be heard for miles--making the woods ring again. So Mr. Martin describes the cry of the agile Gibbon as "overpowering and deafening" in a room, and "from its strength, well calculated for resounding through the vast forests." Mr. Waterhouse, an accomplished musician as well as zoologist, says, "The Gibbon's voice is certainly much more powerful than that of any singer I have ever heard." And yet it is to be recollected that this animal is not half the height of, and far less bulky in proportion than, a man.
There is good testimony that various species of Gibbon readily take to the erect posture. Mr. George Bennett, a very excellent observer, in describing the habits of a male 'Hylobates syndactylus' which remained for some time in his possession, says: "He invariably walks in the erect posture when on a level surface; and then the arms either hang down, enabling him to assist himself with his knuckles; or what is more usual, he keeps his arms uplifted in nearly an erect position, with the hands pendent ready to seize a rope, and climb up on the approach of danger or on the obtrusion of strangers. He walks rather quick in the erect posture, but with a waddling gait, and is soon run down if, whilst pursued, he has no opportunity of escaping by climbing.... When he walks in the erect posture he turns the leg and foot outwards, which occasions him to have a waddling gait and to seem bow-legged."
Dr. Burrough states of another Gibbon, the Horlack or Hooluk: "They walk erect; and when placed on the floor, or in an open field, balance themselves very prettily, by raising their hands over their head and slightly bending the arm at the wrist and elbow, and then run tolerably fast, rocking from side to side; and, if urged to greater speed, they let fall their hands to the ground, and assist themselves forward, rather jumping than running, still keeping the body, however, nearly erect."
Somewhat different evidence, however, is given by Dr. Winslow Lewis:
"Their only manner of walking was on their posterior or inferior extremities, the others being raised upwards to preserve their equilibrium, as rope-dancers are assisted by long poles at fairs. Their progression was not by placing one foot before the other, but by simultaneously using both, as in jumping." Dr. Salomon Muller also states that the Gibbons progress along the ground by a short series of tottering jumps, effected only by the hind limbs, the body being held altogether upright.
But Mr. Martin (l. c. p. 418), who also speaks from direct observation, says of the Gibbons generally:
"Pre-eminently qualified for arboreal habits, and displaying among the branches amazing activity, the Gibbons are not so awkward or embarrassed on a level surface as might be imagined. They walk erect, with a waddling or unsteady gait, but at a quick pace; the equilibrium of the body requiring to be kept up, either by touching the ground with the knuckles, first on one side then on the other, or by uplifting the arms so as to poise it. As with the Chimpanzee, the whole of the narrow, long sole of the foot is placed upon the ground at once and raised at once, without any elasticity of step."
After this mass of concurrent and independent testimony, it cannot reasonably be doubted that the Gibbons commonly and habitually assume the erect attitude.
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