Part 45

Cuvier remarks, that the palm of the fossil horn increases in breadth as it extends outwardly, while that of the elk is broadest next the beam.

The palm of the elk’s horn is directed more backwards, while the fossil one extends more in the lateral direction. The antlers of the elk are shorter and more numerous than those of the fossil animals.

As the horns of the fossil animal exceed in size those of the elk, so, on the contrary, does the skull of the latter exceed in size that of the former; the largest heads of the fossil species not exceeding one foot nine inches in length, while the head of the elk is frequently two feet. The fossil head is broader in proportion; its length being to its breadth as two to one; in the elk they are as three to one, according to Parkinson. The breadth of the skull between the roots of the horns is but four inches in the fossil skulls; in that of the elk in the Society’s Museum it is 6½ inches.

Cuvier thinks it probable that the females of the fossil species had horns, an opinion to which I am very much disposed to subscribe, from having observed that these parts present differences in size and strength, which appear not to be dependent on differences of age. For instance, the teeth of the specimen in Trinity College are much more worn down, and the sutures of the skull are more effaced than in the specimen described in this paper; yet the horns of the latter are much more concave, and more expanded, than those of the former; and on comparing a single horn of each of these specimens together, that belonging to the Society exceeds the other by nearly a sixth in the length, and little less than a third in the breadth; it is not, therefore, unlikely that the animal whose horns were larger and more curved was a male. Something similar to this is observed in the rein-deer, both sexes of which have horns, but with this difference, that they are smaller and less branched in the female. Hence we find that this animal possessed characters of its own sufficient to prove it of a species as distinct from the moose or elk as this latter species is from the rein-deer or any other. Therefore, it is improper to retain the name of elk or moose deer any longer: perhaps it might be better called the Cervus megaceros, a name merely expressive of the great size of its horns.

That this animal shed its head furniture periodically, is proved by the occasional occurrence of detached horns having the smooth convex surface below the burr, similar to what is observed on the cast horns of all deer. Specimens of this are to be seen in the Museum of Trinity College, and I possess one myself, of which I have had a drawing made. As every other species of deer shed their horns annually, there is no reason for supposing that that process occurred at longer intervals in this.

It is a popular opinion with the Indians that the elk is subject to epilepsy, with which he is frequently seized when pursued, and thus rendered an easy prey to the hunters. Many naturalists affect to disbelieve this account, without, however, assigning any sufficient reason. But if it be considered, that, during the growth of the horns, there must be a great increased determination of blood to those parts, which are supplied by the frontal artery, a branch from the internal carotid, it is quite conformable to well established pathological principles, to suppose, that, after the horns are perfected, and have ceased to receive any more blood, that fluid may be determined to those internal branches of the carotid which supply the brain, and establish a predisposition to such derangements of its circulation as would produce epilepsy, or even apoplexy: if such an effect were produced in consequence of the size of the horns in the elk, it is reasonable to suppose that it prevailed in a greater degree in the fossil animal whose horns were so much larger.

What could have been the use of these immense horns? It is quite evident that they would prevent the animal making any progress through a thickly wooded country, and that the long, tapering, pointed antlers were totally unfit for lopping off the branches of trees, a use to which the elk sometimes applies his horns, and for which they seem well calculated, by having their antlers short and strong, and set along the edge of the palm, somewhat resembling the teeth of a saw in their arrangement. It would rather appear, then, that they were given the animal as weapons for its protection, a purpose for which they seem to have been admirably designed; for their lateral expansion is such, that should occasion require the animal to use them in his defence, their extreme tips would easily reach beyond the remotest parts of his body; and if we consider the powerful muscles for moving the head, whose attachments occupied the extensive surfaces of the cervical vertebræ, with the length of the lever afforded by the horns themselves, we can easily conceive how he could wield them with a force and velocity which would deal destruction to any enemy having the hardihood to venture within their range.

From the formidable appearance of these horns, then, we must suppose that their possessor was obnoxious to the aggressions of some carnivorous animals of ferocious habits; and such we know to have abounded in Ireland, as the wolf, and the celebrated Irish wolf dog. Nor would it be surprising if limestone caves should be discovered in this country, containing the remains of beasts of prey and their victims, similar to the hyænas’ dens of Kirkdale, and other places, respecting which such interesting researches have been lately laid before the public by the geologists of this country and the Continent.

The absence of all record, or even tradition, respecting this animal, naturally leads one to inquire whether man inhabited this country during its existence? I think there is presumptive evidence in the affirmative of this question, afforded by the following circumstances. A head of this animal described by Professor Goldfuss of Bonn, was discovered in Germany in the same drain with several urns and stone hatchets; and in the 7th volume of the Archæologia Britannica, is a letter of the Countess of Moira, giving an account of a human body found in gravel, under eleven feet of peat soaked in the bog water: it was in good preservation, and completely clothed in antique garments of hair, which her ladyship thinks might have been that of our fossil animal. But more conclusive evidence on this question is derived from the appearance exhibited by a rib, presented by Archdeacon Maunsell to the Royal Dublin Society, in which I discovered an oval opening near its lower edge, the long diameter of which is parallel to the length of the rib, its margin is depressed on the outer, and raised on the inner surface, round which there is an irregular effusion of callus. This opening had been evidently produced by a sharp pointed instrument, which did not penetrate so deep as to cause the animal’s death, but which remained fixed in the opening for some length of time afterward; in fact it was such an effect as would be produced by the head of an arrow remaining in a wound after the shaft was broken off.

It is not improbable, therefore, that the chace of this gigantic animal once supplied the inhabitants of this country with food and clothing.

As to the causes which led to the extinction of this animal, whether it was suddenly destroyed by the deluge, or by some other great catastrophe of nature, or whether it was ultimately exterminated by the continued and successful persecution of its pursuers, as has nearly been the case with the red deer within the recollection of many of the present generation, I profess myself unable to form any decided opinion, owing to the limited number of facts as yet collected on the subject. On some future occasion I may, perhaps, be induced to revert to so interesting a topic, should I have opportunities of discovering any thing worthy of communication.

The following Table exhibits a comparative view of the measurements of different parts of the skeletons of the Cervus Megaceros in the Museum of the Royal Dublin Society, and in the Royal Museum of the University of Edinburgh, with some parts of the Moose. The measurements of the Edinburgh specimen are taken from Professor Jameson’s memoir on organic remains, in the Supplement to the Encyclopedia Britannica.

|R. D. Soc. |U. of Edin.| Moose HEAD. | Ft. In. | Ft. In. | Ft. In. | | | Length of the head, | 1 8½ | 1 8¼ | Breadth of the skull between | | | the orbits. | 0 10½ | 0 9 | Do. of skull at the occiput, | 0 8 | | Diameter of the orbit, | 0 2⅜ | 0 2½ | Distance between infra orbitar | | | holes across the skull, | 0 7 | | Length of alveolar processes | | | of the upper jaw, | 0 6 | 0 6 | Length of lower jaw, | 1 5½ | 0 3½ | Diam. of foramen magnum, | 0 2 | | | | | HORNS. | | | | | | Distance between the extreme | | | tips, measured by the skull, | 11 10 | | Ditto, in a straight line | | | across, | 9 2 | 6 8 | 3 7 Length of each horn, | 5 9 | 5 1 | Greatest breadth of the palm, | 2 10 | | Length of the beam, | 1 9 | | 0 6½ Ditto of brow antler, | 0 8¾ | | Ditto of sur-antler, | 1 4 | | Circumference of the beam | | | at root of brow antler, | 1 0¾ | | 0 7½ | | | BODY. | | | | | | Length of spine, | 10 10 | 9 8 | Ditto of sternum, | 2 4 | | Height to the upper extremity | | | of the dorsal spines, | 6 6 | | Ditto to the highest point | | | of the tip of the horn, | 10 4 | | | | | EXTREMITIES. | | | | | | Greatest length of the scapula,| 1 6½ | | Ditto breadth at the base, | 0 10¾ | | Ditto depth of its spine, | 0 2¾ | | Length of the humerus, | 1 4 | 1 3½ | Ditto of ulna and radius, | 1 8 | 1 6 | Ditto of carpus, | 0 2¾ | 0 2 | Circumference of do., | 0 9½ | | Length of metacarpus, | 1 0½ | 1 0½ | Length of phalanges, | 0 7 | 0 6½ | From anterior superior spine | | | of one ileum to that of | | | the other, | 1 4½ | 1 6½ | From anterior superior spine | | | to the tuber ischii, | 1 8 | 1 9½ | Greatest diameter of | | | foramen ovale, | 0 4 | 0 3 | Least do. of do., | 0 2¾ | 0 2¼ | Length of the femur, | 1 6½ | 1 5½ | Ditto of tibia, | 1 6 | 1 6 | Length of the tarsus, | | | including the os calcis, | 0 8 | | Ditto of the metatarsus, | 1 1¾ | 1 1¾ |

2. Account of the Two Living Species of Elephant, and of the Extinct Species of Elephant, or Mammoth.

1. ELEPHAS AFRICANUS.--The Elephant with rounded skull, large ears, grinders, having rhomboidal-shaped marks on their crown, which we call the African Elephant (Elephas Africanus), is a quadruped which has hitherto been found only inhabiting Africa. There can be no doubt that it is this species which lives at the Cape, at Senegal, and in Guinea; there is reason to believe that it also occurs at Mosambique; but it is not certain that individuals of the following species do not occur in this part of Africa. A sufficient number of individuals have not been figured or compared, to know if this species presents remarkable varieties. It is it that produces the largest tusks. Both sexes are equally furnished with tusks, at least at Senegal. The natural number of the hoofs is four before, and three behind. The ear is very large, and covers the shoulder. The skin is of a deep and uniform brown. This species has not been domesticated in modern times. It appears, however, to have been tamed by the ancients, who attributed to it less power and courage in that state than to the following species; but their observations do not appear to have been confirmed, at least in so far as refers to magnitude. Its natural manners are not perfectly known; yet judging of them by the notices of travellers, they appear to resemble in every thing essential those of the following species.

2. ELEPHAS INDICUS.--The Elephant with elongated skull, concave forehead, small ears, grinders marked with undulating bands, which we call the Indian Elephant (Elephas Indicus), is a quadruped which has only been observed with certainty beyond the Indus. It extends from both sides of the Ganges to the Eastern Sea and the south of China. They are also found in the Islands of the Indian Sea, in Ceylon, Java, Borneo, Sumatra, &c. There is still no authentic proof that it exists in any part of Africa, although neither is the contrary absolutely proved. The inhabitants of India having from time immemorial been in the habit of taking this species and taming it, it has been much better observed than the other. Varieties have been remarked as to size, lightness of form, the length and direction of the tusks, and the colours of the skin. The females and some of the males have tusks which are always small and straight. The tusks of the other males never attain so great a length as in the African species. The natural number of the hoofs is five before and four behind. The ear is small, frequently angular. The skin is commonly grey, spotted with brown. There are individuals entirely white. The height varies from fifteen to sixteen feet. Its manners, the mode of taking it, and of treating it, have been carefully described by many travellers and naturalists, from Aristotle down to Mr Corse Scott.

3. ELEPHAS PRIMIGENIUS, Blum, or MAMMOTH.--The Elephant with elongated skull, concave forehead, very long alveolæ for the tusks, the lower jaw obtuse, the grinders broader, parallel, marked with closer bands, which we name the Fossil Elephant (Elephas primigenius, Blum.), is the Mammoth of the Russians. Its bones are only found in the fossil state. No person has seen in a fresh state bones resembling those by which this species is peculiarly distinguished, nor have the bones of the two preceding species been seen in the fossil state. Its bones are found in great number in many countries, but in better preservation in the north than elsewhere. It resembles the Indian more than the African species. It differs, however, from the former in the grinders, in the form of the lower jaw, and many other bones, but especially in the length of the alveolæ and tusks. This last character must have singularly modified the figure and organisation of its proboscis, and given it a physiognomy much more different from that of the Indian species, than might have been expected from the similarity of the rest of their bones. It appears that its tusks were generally large, frequently more or less spirally arcuate, and directed outwards. There is no proof that they differ much according to differences of sex or race. The size was not much greater than that to which the Indian species may attain; it appears to have been still clumsier in its proportions. It is already manifest from its osseous remains, that it was a species differing more from the Indian, than the ass from the horse, and the jackal and isatis from the wolf and fox. It is not known what had been the size of its ears, or the colour of its skin; but it is certain that, at least, some individuals bore two sorts of hair, namely, a red, coarse, tufted wool, and stiff black hairs, which, upon the neck and along the dorsal spine, became long enough to form a sort of mane. Thus, not only is there nothing impossible in its having been able to support a climate which would destroy the Indian species, but it is even probable that it was so constituted as to prefer cold climates. Its bones are generally found in the alluvial and superficial strata of the earth, and most commonly in the deposits which fill up the bottom of valleys, or which border the beds of rivers. They scarcely ever occur by themselves, but are confusedly mingled with bones of other quadrupeds of known genera, such as rhinoceroses, oxen, antelopes, horses, and frequently with remains of marine animals, particularly conchiferous species, some of which have even been found adhering to them. The positive testimony of Pallas, Fortis, and many others, does not allow us to doubt that this latter circumstance has frequently taken place, although it is not always observed. We ourselves have at this moment under our eyes a portion of a jaw covered with millepores and small oysters.

The strata which cover the bones of elephants are not of very great thickness, and they are scarcely ever of a rocky nature. They are seldom petrified, and there are only one or two cases recorded in which they were found imbedded in a shelly or other rock. Frequently they are simply accompanied with our common fresh water shells. The resemblance, in this latter respect, as well as with regard to the nature of the soil, between the three places, of which we have the most detailed accounts, viz. Tonna, Cantstadt, and the Forest of Bondi, is very remarkable. Every thing, therefore, seems to announce that the cause which has buried them, is one of the most recent of those that have contributed to change the surface of the globe. It is nevertheless a physical and general cause; the bones of fossil elephants are so numerous, and have been found in places so desert and even uninhabitable, that we cannot suppose that they had been conducted there by man. The strata which contain them and those which are above them, shew, that this cause was aqueous, or that it was water that covered them; and in many places these waters were nearly the same as those of our present sea, since they supported animals nearly the same. But, it was not by these waters that they were transported to the places where they now are. Bones of this description have been found in almost every country that has been examined by naturalists. An irruption of the sea that might have brought them from places which the Indian elephant now inhabits, could not have scattered them so far, nor dispersed them so equably. Besides, the inundation which buried them has not risen above the great chains of mountains, since the strata which it has deposited, and which cover the bones, are only found in plains of little elevation. It is not, therefore, seen how the carcases of elephants could have been transported into the north, across the mountains of Thibet, and the Altaic and Uralian chains.

Further, these bones are not rolled; they retain their ridges and apophyses; they have not been worn by friction. Very frequently the epiphyses of those which had not yet attained their full growth, are still attached to them, although the slightest effort would suffice to detach them. The only alterations that are remarked, arise from the decomposition which they have undergone during their abode in the earth. Nor can it with more reason be represented that the entire carcases had been violently transported. In this case, the bones would indeed have remained entire; but they would also have remained together, and would not have been scattered. The shells, millepores, and other marine productions which are attached to some of these bones, prove besides that they had remained at least some time stripped and separated at the bottom of the fluid which covered them. The elephants’ bones had therefore already been in the places in which they are found, when the fluid covered them. They were scattered about in the same manner as in our own country the bones of horses and other animals that inhabit it may be, and as the dead bodies are spread in the fields.

Every circumstance, therefore, renders it extremely probable, that the elephants which have furnished the fossil bones, dwelt and lived in the countries where their bones are at present found. They could only, therefore, have disappeared by a revolution, which had destroyed all the individuals then living, or by a change of climate, which prevented them from propagating. But whatever this cause may have been, it must have been sudden. The bones and ivory which are found in so perfect a state of preservation in the plains of Siberia, are only so preserved by the cold which congeals them there, or which, in general, arrests the action of the elements upon them. If this cold had come on by degrees and slowly, these bones, and still more the soft parts with which they are still sometimes invested, would have had time to decompose, like those which occur in warm and temperate countries. It would especially have been impossible that an entire carcase, like that discovered by Mr Adams, could have retained its flesh and skin without corruption, if it had not been immediately enveloped by the ice which preserved it. Thus, all the hypotheses of a gradual cooling of the earth, or of a slow variation, whether in the inclination or in the position of the axis of the globe, fall to be rejected.

If the present elephants of India were the descendants of these ancient elephants, which have been preserved in that climate to the present day, from their being there placed beyond the reach of the catastrophe which destroyed them in the others, it would be impossible to explain why their species has been destroyed in America, where remains are still found, which prove that they had formerly existed there. The vast empire of Mexico presented to them heights enough to escape from an inundation so little elevated as that which we must suppose to have taken place, and the climate there is warmer than is requisite for their temperament.

The various mastodons, the hippopotamus and the fossil rhinoceros lived in the same countries, and in the same districts, as the elephants, since their bones are found in the same strata and in the same state. Yet these animals very assuredly no longer exist. Every thing therefore, Cuvier maintains, concurs to induce a belief that the fossil elephant is, like them, an extinct species, although it resembles more than they one of the species at present existing, and that its extinction has been produced by a sudden cause, by the same great catastrophe which destroyed the species of the same epoch.

3. On the Great Mastodon, or Animal of the Ohio.

It appears that the Great Mastodon or Animal of the Ohio, was very like the elephant in its tusks and whole skeleton, the grinders excepted; that it very probably had a proboscis; that its height did not exceed that of the elephant, but that it was a little more elongated, and had limbs somewhat thicker, with a more slender belly. Notwithstanding all these points of resemblance, the peculiar structure of its grinders is sufficient to constitute it of a different genus from the elephant. It further appears, that it fed much in the same manner as the hippopotamus and boar, choosing by preference the roots and other fleshy parts of vegetables; that this sort of food must have drawn it towards the soft and marshy places; that, nevertheless, it was not formed for swimming, and living often in the water like the hippopotamus, but that it was a true land animal. Its bones are much more common in North America than any where else. They are even perhaps exclusively confined to that country. They are better preserved, and fresher, than any other fossil bones known; and, nevertheless, there is not the slightest proof, the smallest authentic testimony, calculated to impress a belief that either in America, or any where else, there is still any living individual, for the various accounts which we have from time to time read in the journals respecting living mastodons, which had been observed in the forests or plains of that vast continent, have never been confirmed, and can only pass for fables.

NOTE

ON THE CAVES IN WHICH BONES OF CARNIVOROUS ANIMALS OCCUR IN GREAT QUANTITIES.

The extraordinary accumulations of fossil bones in caves and caverns in different districts, especially in those composed of limestone, have for many years engaged the attention of inquirers; and, of late, have afforded many interesting facts to the geologist and zoologist. In England, as will appear from the following details, many different fossil animals have been discovered in limestone caves; but hitherto the caves in Scotland, which will probably be found to contain interesting documents of an ancient population, have not been examined. As the subject is a curious and interesting one, we shall, in the following pages, principally from Cuvier’s great work, lay before our readers a pretty full account of the different caves, especially those that afford bones of carnivorous animals.

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