Part 27

They differ, however, from these latter, in having the fore ones more simple, the backmost of all with three necks, and the upper ones rhomboidal, and marked with ridges very much resembling those of the rhinoceros.

We are still ignorant what the form of their snout, and the number of their toes, may have been. I have discovered not less than twelve species of this genus, all in France, deposited in marly rocks of fresh-water formation, and filled with lymneæ and planorbes, which are shells peculiar to pools and marshes.

The largest species is found near Orleans, in the same quarry as the palæotheria; it approaches the rhinoceros.

There is a smaller species in the same place; a third occurs at Montpellier; a fourth near Laon; two near Buchsweiler in Alsace; five near Argenton in Berry; and one of the three occurs again near Issel, where there are also two others. There is also a large one near Gannat.

These species differ from each other in size, the smallest being scarcely so large as a lamb of three months, and in various circumstances connected with the form of their teeth, which it would be too tedious and minute to detail here.

The Anoplotheria have hitherto been discovered nowhere but in the gypsum quarries of the neighbourhood of Paris. They have two characters which are observed in no other animal; feet with two toes, the metacarpal and metatarsal bones of which are separate in their whole length, and do not unite into a single piece, as in the ruminantia; and teeth placed in a continuous series without any interruption. Man alone has the teeth so placed in mutual contiguity, without any interval. Those of the anaplotheria consist of six incisors in each jaw, a canine tooth and six grinders on each side, both above and below; their canine teeth are short and similar to the outer incisors. The three first grinders are compressed; the four others are, in the upper jaw, square, with transverse ridges, and a small cone between them; and, in the lower jaw, in the form of a double crescent, but without neck at the base. The last has three crescents. Their head is of an oblong form, and does not indicate that the muzzle has terminated either in a proboscis or a snout.

This extraordinary genus, which can be compared to nothing in living nature, is subdivided into three subgenera: the Anaplotheria, properly so called, the anterior molares of which are still pretty thick, and the posterior ones of the lower jaw have their crescents with a simple ridge; the Xiphodons, of which the anterior molares are thin and sharp on the edges, and the under posterior, have, directly opposite the concavity of each of their crescents, a point, which, on being worn, also assumes the form of a crescent, so that then the crescents are double as in the ruminantia; lastly, the Dichobunes, the outer crescents of which are also pointed at the beginning, and which have thus points disposed in pairs upon their lower posterior grinders.

The most common species in our gypsum quarries (An. commune), is an animal of the height of a boar, but much more elongated, and furnished with a very long and very thick tail, so that altogether it has nearly the proportion of the otter, but larger. It is probable that it was well fitted for swimming, and frequented the lakes in the bottom of which its bones have been incrusted by the gypsum which was deposited there. We have one a little smaller, but in other respects pretty similar (An. secundarium.)

We are as yet acquainted with only one xiphodon, which, however, is a very remarkable animal: it is that which I have named An. gracile. It is slender, and delicately formed, like the prettiest gazelle.

There is one dichobune, nearly of the size of a hare, to which I have given the name of An. leporinum. Besides its subgeneric characters, it differs from the anaplotheria and xiphodons, in having two small and slender toes on each foot, at the sides of the two large toes.

We do not know if these lateral toes exist in the two other dichobunes, which are small, and scarcely exceed in size the common Guinea pig.

The genus of Anthracotheria is in some degree intermediate between the palæotheria, anaplotheria, and hogs. I have named it so, because two of its species have been found in the lignites of Cadibona, near Savone. The first approached the rhinoceros in size; the second was much smaller. They have also been found in Alsace, and in the Vélay. Their grinders are similar to those of the anaplotheria; but they have projecting canine teeth.

The genus Cheropotamus is found in our gypsum deposits, where it accompanies the palæotheria and anaplotheria, but where it is of much rarer occurrence. Its posterior grinders are square above, rectangular below, and have four large conical eminences surrounded with smaller ones. The anterior molares are short cones, slightly compressed, and with two roots. Its canine teeth are small. Neither its incisors nor its feet are yet known. I possess only one species, which is of the size of a Siam hog.

The genus Adapis has also but one species, which is at most of the size of a rabbit: it is also from our gypsum quarries, and must have been nearly allied to the anaplotheria.

We have thus nearly forty species of pachydermata belonging to genera now entirely extinct, and presenting forms and proportions to which there is nothing that can be compared in the present animal kingdom, excepting two tapirs and a daman.

This large number of pachydermata is so much the more remarkable, that the ruminantia, which are at present so numerous in the genera of deer and antelopes, and which attain so great a size in those of the oxen, giraffes, and camels, scarcely make their appearance in the deposits of which we are speaking.

I have not seen the slightest trace of them in our gypsum quarries; and all that has come to my hands consists of some fragments of a deer, of the size of the roe, but of a different species, collected among the palæotheria of Orleans; and of one or two other small fragments, from Switzerland, which, however, are perhaps of doubtful origin.

But our pachydermata have not for all this been the only inhabitants of the countries in which they lived. In our gypsum deposits, at least, we find along with them carnivora, glires, several sorts of birds, crocodiles, and tortoises; and these two latter genera also accompany them in the molasse sandstones and marly deposits of the middle and south of France.

At the head of the carnivora, I place a Bat, very recently discovered at Montmartre, and which belongs to the proper genus Vespertilio. The existence of this genus, at an epoch so remote, is so much the more surprising, that, neither in this formation, nor in those which have succeeded it, have I seen any other trace, either of cheiroptera or of quadrumana: no bone or tooth of either monkey or maki has ever presented itself to me, in the course of my long researches.

Montmartre has also furnished the bones of a fox different from ours, and which also differs from the jackals, isatises, and the various species of foxes peculiar to America; those of a carnivorous animal allied to the raccoons and coaties, but larger than any known species; those of a particular species of civet; and of two or three other carnivora, which it has not been possible to determine, from the want of tolerably complete portions.

What is still more remarkable, is, that there are skeletons of a small sarigue, allied to the marmose, but different, and consequently of an animal belonging to a genus which is at the present day confined to the New World. Skeletons of two small glires, of the genus myoxus, and a skull belonging to the genus sciurus, have also been collected.

Our gypsum deposits are more fertile in bones of birds than any of the other strata either anterior or posterior to it. Entire skeletons, and parts of at least ten species belonging to all the orders, are found there.

The crocodiles of the period in question approach our common crocodiles in the form of the head, while, in the deposits of the Jura period, we find only species allied to the gavial.

A species has been found at Argenton, which is remarkable for its compressed, sharp teeth, having their edges dentated like those of certain monitors. Some remains of it also occur in our gypsum quarries.

The tortoises of this period are all fresh-water ones: some of them belong to the subgenus Emys; and there are species, both at Montmartre, and still more especially in the molasse sandstones of the Dordogne, which are larger than any living species known; the others are Trionyces or soft tortoises. This genus, which is easily distinguished by the vermiculate surface of the bones of its shell, and which at present exists only in the rivers of warm countries, such as the Nile, the Ganges, and the Orinoko, has been very abundant in the places where the palæotheria lived. Vast quantities of its remains are found at Montmartre, and in the molasse sandstones of the Dordogne, and the other gravel deposits of the south of France.

The fresh-water lakes, around which these various animals have lived, and which had received their bones, nourished, besides the tortoises and crocodiles, some fishes and testaceous mollusca. All that have been collected of these two classes of animals, are as foreign to our climate, and even as much unknown in our present waters, as the palæotheria, and other quadrupeds which were coeval with them.

The fishes have even in part belonged to unknown genera.

Hence, it cannot be doubted that this race of inhabitants, which might be termed the population of the middle age, this first great production of mammifera, has been entirely destroyed; and, in fact, in all places where remains of them have been discovered, there are great deposits of marine formation above them, so that the sea has overwhelmed the countries which these races inhabited, and has rested upon them during a long period of time.