Part 76

Skull and teeth.--Adult male: See measurements and plates 22-24. As described in Mustela frenata frenata except that: Weight, 8 grams; basilar length 54; least width of palate less than length of P4; anterior margin of masseteric fossa anterior to middle of m2.

Female: Skull unknown.

Remarks.--The type specimen, taken by the veteran collector of Texan mammals, H. P. Attwater, appears to have been the first one of these animals to find its way into the collection of any museum or naturalist. The second specimen from Kerr County was secured by, or through, the well-known commercial collector, F. B. Armstrong. Two trade skins, from Kerr County, taken on December 10, 1938, are in the Texas Cooperative Research Collection, as is also the skeleton of a young animal from Fredericksburg. The two other specimens from McLennan County (both males contrary to the statement of Strecker, 1924:14), owe their preservation to the alertness of John K. Strecker, Curator of the Baylor University Museum, who has given a complete account of their history.

The range of this subspecies is thought to include much of central Texas.

The preserved parts of the skin of the type specimen show it to have been generally large. The part of the tail preserved measures 226 millimeters and the skin of the head and neck is correspondingly large. The skin alone, no. 427, from near Waco, Texas, has, as now stuffed, a body 365 millimeters long. Individuals of this race attain larger size than those of any other American member of the subgenus Mustela with the possible exception of Mustela frenata macrophonius from Veracruz, México. In addition to large size, texensis and macrophonius are analogous in that each has a small geographic range at the northern end of an extensive range of its similarly colored southern relative from which it differs mainly in size. Each of the two groups, goldmani and macrophonius on the one hand and perotae, frenata and texensis on the other, has relatively uniform color, color pattern and body proportions over a large region but at its northern extremity develops a "giant" population, M. f. macrophonius and M. f. texensis, respectively. The skull of the type specimen of M. f. texensis is the largest one seen of any American weasel. The type specimen of M. f. macrophonius has a basilar length that is greater by one-tenth of a millimeter but in every other measurement taken the skull of M. f. texensis is the larger. Its weight, 8 grams, also shows it to be larger.

The broad, white bands in front of the ears are confluent with the white patch between the eyes on both sides in two specimens and on one side only in one other specimen. A white patch between the ears is present in four specimens. The dark spot at each angle of the mouth is absent on both sides in four specimens and on one side only in one other specimen. Thus out of a possible twelve cases, the broad bands in front of the ears are confluent with the spot between the eyes in five cases. Four of the six specimens have a white spot between the ears. The dark spot at each angle of the mouth is present three out of a possible twelve times.

The skull of no. 2017, from five miles north of Waco, is smaller than either of the two skulls seen from Kerr County and in this respect approaches M. f. frenata. There is no actual evidence of intergradation with any other subspecies but intergradation probably does take place with M. f. neomexicana and possibly with M. f. arthuri and M. f. primulina.

Strecker (1924:14) remarks that of the two specimens obtained near Waco, one was taken in a trap baited for mink and the other was shot in a hen house. None of the four skulls had the frontal sinuses infested with parasites.

Specimens examined.--Total number, 7, arranged by counties from north to south.

=Texas.= McLennan County: 5 mi. N Waco, 1; Erath, 1. Gillespie County: Fredericksburg, 1. Kerr County: 4; 1; and 2 trade skins.

=Mustela frenata frenata= Lichtenstein

Long-tailed Weasel

Plates 1, 22, 23, 24, 36, 37, 38 and 40

Mustela frenata Lichtenstein, Darstellung neuer oder wenig bekannter Säugethier, 1832, pl. 42, and corresponding text, unpaged; Seton, Lives of game animals, 2:576, 1929.

Mustela brasiliensis Sevastianoff, Mem. de l'Acad. Imp. Sci. St. Petersburg, 4:356-363, tab. 4, 1813, name on plate only, the description being in the text (not of Gmelin, 1788); Gray, Proc. Zoöl. Soc. London, 1865:114.

Putorius frenatus, Baird Mamms. N. Amer., p. 173, 1858; Merriam, N. Amer. Fauna, 11:26, pl. 3, figs. 1, 1a, 1b, June 30, 1896; Bailey, N. Amer. Fauna, 25:198, October 24, 1905.

Putorius (Gale) brasiliensis aequatorialis Coues, Fur-bearing animals, p. 142, 1877, part? ("merely as a substitute for Gray's [supposedly] preoccupied name" that is, aureoventris).

Putorius (Gale) brasiliensis frenatus, Coues, Fur-bearing animals, p. 142, 1877 (part).

Putorius mexicanus Coues, Fur-bearing animals, p. 142, 1877, [nomen nudum, cited by Coues in synonymy as "Putorius mexicanus, Berlandier, MSS. ic. ined. 4 (Tamaulipas and Matamoros)"].

Putorius brasiliensis frenata, Allen, Bull. Amer. Mus. Nat. Hist., 3:219, April 17, 1891.

Putorius brasiliensis frenatus, Allen, Bull. Amer. Mus. Nat. Hist., 6:197, May 31, 1894; Bangs, Proc. Biol. Soc. Washington, 10:9, February 25, 1896; Allen, Bull. Amer. Mus. Nat. Hist., 8:74, April 22, 1896.

Mustela frenata frenata, Strecker, The Baylor Bull., 27:12, August, 1926; Hall, Carnegie Instit. Washington Publ. 473:108, November 20, 1936.

Type.--Female, adult, skull and skin; no. 991, Berlin Zool. Mus., México City, México; June, 1829; obtained by F. Deppe.

The specimen once mounted, now is remade into a study skin and lacks the distal part of the tail. The skull (plates 36-38, 40) lacks the basicranial region.

Range.--Altitudinally, sea level (Brownsville, Texas) to 7600 feet (Tlalpam, México); from southern Texas as far south as México City; Lower Sonoran to at least Transition life-zone. See figure 29 on page 221.

Characters for ready recognition.--Differs from M. f. perotae in nonextension of blackish over anterior fourth of neck, least width of color of underparts more than 37 per cent of greatest width of color of upper parts; height of tympanic bulla more than distance from its anterior margin to foramen ovale; from M. f. leucoparia by restricted white facial markings that cover less than half surface of head in front of ears, by nonextension of black of head onto anterior half of neck and by wider (more than 7.8) tympanic bullae; from M. f. neomexicana by Brussels Brown rather than Buckthorn Brown color of upper parts and mastoid breadth less than postpalatal length; from M. f. texensis by smaller size of body and skull (basilar length in adult males less than 52.5); from M. f. arthuri by white facial markings and postorbital breadth less than distance between posterior borders of P4 and P2; from M. f. tropicalis by nonextension of blackish over anterior fourth of neck, least width of underparts more than 37 per cent of greatest width of upper parts, postorbital breadth of adult males less than distance between posterior borders of P4 and P2.

Description.--Size.--Male: Fifteen adults and subadults from Brownsville, Texas, yield average and extreme measurements as follows: Total length, 485 (430-556); length of tail, 202 (165-250); length of hind foot, 48 (40-55). Averages believed to be reliable but extremes probably are not. Tail averages 71 per cent as long as head and body. Length of hind foot less than basal length. Corresponding measurements of an adult male (topotype, no. 50826) from Tlalpam, México, are: 505, 203, 53. Another adult male, from Miquihuana, Tamaulipas, México, measures: 520, 215, 52.

Female: Six adults, subadults and young from Brownsville, Texas, yield average and extreme measurements as follows: Total length, 420 (362-456); length of tail, 173 (126-200); length of hind foot, 41 (40-46). Tail averages 70 per cent as long as head and body. Length of hind foot more (with possible exception of no. 36362/48732 U. S. Nat. Mus.) than basal length.